News 

13th International Congress of Auchenorrhyncha
2010-08-16
Held in Vaison-la-Romaine, France (28 June-2 July 2010), the Congress included a presentation by Olivia Evangelista, D. M. Takiya, and A. M. Sakakibara: “Reassessment of the treehopper tribe Procyrtini: morphological variation and phylogenetic placement based on 28S rDNA (Membracidae: Darninae)” and a poster by Stuart H. McKamey: “The first immatures of two plesiomorphic genera of Membracidae and their Phylogenetic implications,” which focused on Antillotolania and Deiroderes. The Congress website includes details of the overall program. [S. H. McKamey and L. L. Deitz]

Recent Treehopper Publications
2010-08-16
Please contact us to add or correct references.

Hamilton, K. G. A.; Cocroft, R. B. 2010a. Establishing the identity of existing name in the North American Enchenopa binotata species complex of treehoppers (Hemiptera: Membracidae). Entomological News 120(5): 534-565. [designates a female specimen as neotype of both E. binotata and E. porrecta; notes on E. brevis, E. sericea, and undescribed species]

Sakakibara, A. M.; Lencioni-Neto, F. 2009a. Duas espécies novas de membracídeos (Hemiptera, Membracidae) da Mata Atlântica do Estado de São Paulo, Brasil. [Two new species of treehopper (Hemiptera, Membradidae) from the Atlantic Forest of São Paulo, Brazil.]. Papeis Avulsos de Zoologia 49(16): 191-196. [in Portuguese, with English summary; Antonae brasiliensis and Hypheodana gargionei, n. spp.]

Wallace, M. S.; Bartlett, C. R.; Deitz, L. L.; Rothschild, M. J. 2009a. New state records of treehoppers (Hemiptera: Membracidae) for Delaware, Maryland, Mississippi, Pennsylvania, and West Virginia. Proc. Entomol. Soc. Washington 111(3): 734-738. [presents new state records for 27 spp.]

Wallace, M. S.; Maloney, S. M. 2010a. Treehopper (Hemiptera: Membracidae) biodiversity and seasonal abundance in the Pocono till barrens, Long Pond, Pennsylvania. Proc. Entomol. Soc. Washington 112(2): 281-294. [lists 31 spp.; 1 new state record (Cyrtolobus inermis)]

Treehopper Bulletins at Bargain Prices
2010-08-16
Limited numbers of five North Carolina Agricultural Research Technical Bulletins on treehoppers remain available at great prices. Additional shipping fees are charged for overseas orders (prices below include shipping for domestic orders). Payment must be in U.S. dollars (check or money order payable to N.C. State University). Contact: Jeanne Marie Wallace

TB225 $2.50 Classification of the higher categories of the New World treehoppers (Homoptera: Membracidae). Deitz 1975

TB284 $4.00 Bibliography of the Membracoidea (Homoptera: Aetalionidae, Biturritiidae, Membracidae, and Nicomiidae), 1956-1980. Deitz and Kopp 1987

TB290 $3.00 Bibliography of the Membracoidea (Homoptera: Aetalionidae, Biturritiidae, Membracidae, and Nicomiidae), 1981-1987. Deitz 1989

TB293 $10.00 Revision of the Neotropical treehopper tribe Aconophorini (Homoptera: Membracidae). Dietrich and Deitz 1991

TB294 $7.00 Revision of the Neotropical treehopper genus Metcalfiella (Homoptera: Membracidae). McKamey and Deitz 1991

Alfred Keller’s Model of Bocydium globulare
2010-12-13
"The surreal treehoppers," a short note about this artist and treehoppers, appears at the blog: Why Evolution is True. The images are astonishing.

Browse NCSU’s Treehopper Collection through GigaPans
2010-12-13
Through the magic of GigaPans (online, high resolution digital images), one can now browse the drawers of treehoppers and other insects held at the North Carolina State University Insect Museum, NCSU, Raleigh. Funding for this project was provided to Andy Deans and Brian Wiegmann by the National Science Foundation (DBI-0847924). The collection includes Z. P. Metcalf’s specimens and many more recent additions.

Aetalionidae

Melizoderidae

Membracidae

18th Treehopper Gathering Enjoys “Aliens of the Amazon: Treehoppers
2010-12-13
The 2010 Treehopper Gathering was held at the Little Orleans Campground, Little Orleans, Maryland, 4-6 June 2010, with Charles Bartlett (University of Delaware) serving as the organizer. This annual event, initiated by Tom Wood, first met in the spring of 1989 at the University of Delaware. Nearly 25 entomologists and students, mostly from institutions in the eastern U.S. (Delaware, Kansas, Maryland, New York, North Carolina, Pennsylvania, South Dakota, and Washington, D.C.) collected treehoppers and other insects, worked on collaborative projects, and enjoyed camping, eating and swimming. After a fantastic feast of turkey and steak, Stuart McKamey (Systematic Entomology Laboratory, National Museum of Natural History, Washington, D.C. ) shared a TV documentary featuring treehoppers filmed in an expedition to Ecuador (November 2008). The venture, financed by Mona Lisa Productions (Lyon France), included Stuart, who collaborated in organizing the trip, as well as membracid researchers Rex Cocroft (University of Missouri ) and Carolina Godoy (University of Costa Rica). The result is an award-winning documentary: “Aliens of the Amazon: Treehoppers,” which has been released in various versions in the U.S., Canada, France, Germany, Austria, Italy, and Belgium. [L. L. Deitz, M. S. Wallace, and S. H. McKamey]


Most Wanted List
2011-01-16
We invite photographers to submit original high quality digital images of treehoppers for possible inclusion on our website through contact us. Please include copyright information (holder and year), name of taxon (if known), and other relevant data for each image. Selected images are featured on the "Top Pics" page.

Our current most wanted images include the genera Acanthicoides, Aleptocentrus, Ananthasubramanianum, Aspasiana, Centrobelus, Demanga, Gargarina, Gonoconophora, Insitor, Insitoroides, Matonotus, Megaloschema, Neocentrus, Neomachaerotypus, Occator, Spathenotus, and Tremembaetalion.

Olivia Evangelista and Stuart McKamey recently provided images of three genera previously on our "Most Wanted List."

2011 Annual Treehopper Gathering: 3-5 June, Little Orleans, Maryland
2011-03-20
The 19th Annual Treehopper Gathering is scheduled for 3-5 June 2011 (arrive Friday, depart Sunday), at the Little Orleans Campground, Little Orleans, Maryland. This informal event provides a great opportunity to talk and collect with others passionate about treehoppers and related insects.

To register or obtain further information, please contact Charles Bartlett. Campsites are somewhat limited, so please register with Charles if you wish to camp with our group. Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening.

Treehoppers Top the Headlines
2011-05-06
An article featured on the cover of Nature confirms what many of us already knew: Treehoppers are Extraordinary!

Prud'homme et al. (2011) present intriguing evidence that the "pronotal" modifications of treehoppers may arise from fused appendages of the prothorax, which are serially homologous with the wings of the meso- and metathorax. Having fused into prothoracic "helmets" with no function in flight, the presumptive wing elements diversified to produce the dazzling regalia that adorn members of the families Membracidae, Aetalionidae, and Melizoderidae. See also: Discover Magazine, Science Now, Why Evolution is True (Video), and Why Evolution is True (Origin of Treehopper Helmet).

Amazing treehoppers also appear in a note on Myrmecomorphs.

[L. L. Deitz]

19th Annual Treehopper Gathering: 3-5 June 2011, Little Orleans, Maryland
2011-08-02
The 2011 Annual Treehopper Gathering included workers interested in treehoppers (and other less outrageous insect groups) from across several eastern states of the US, as well as Kansas and even Brazil. As usual, the camaraderie, collecting, and food were terrific. Institutions represented include: ESU, East Stroudsburg University; FAC, Florida Arthropod Collection; KU, Kansas University Natural History Museum, University of Kansas; MC, Metropolitan College of New York; NCSU, North Carolina State University; NYSM, New York State Museum; SEL, Systematic Entomology Laboratory, USDA, c/ Smithsonian Institution; UD, University of Delaware; and UFPR, Universidade Federal do Paraná, Curitiba-PR, Brazil.



Participants (from left to right): 1, Charles Bartlett, organizer of the event (UD); 2, Cyrus Kasaaian (UD), 3, Ashley Kennedy (UD); 4, Julie Urban (NYSM); 5, Andrew Short (KU); 6, Jeremy Unseld (KU); 7, Crystal Maier (KU); 8, Clay McIntosch (KU); 9, Ruth Thompson (MC); 10, Vinton Thompson (MC); 11, Olivia Evangelista (NYSM & UFPR); 12, Keith Bayless (NCSU);13, Mark Rothschild (FAC); 14, Adam Bell (NYSM); 15, Andressa Paladini (NYSM); 16, Lawrence Barringer (UD); 17, Jessica Newbern (ESU) with 18, Reese Newbern, and 19, Aidan Uher; 20, Matthew Wallace (ESU); 21, Stuart McKamey (SEL); 22 Lewis Deitz (NCSU); 23, Anthony Gonzon (UD); 24, Nadine Jahn (ESU); 25, David Miracle (ESU).



Above, Matt Wallace searches for treehoppers near St. Patricks Church, Little Orleans, Maryland. Fifty-six treehopper species (23 genera) are recorded from Little Orleans (Bartlett et al. 2008, Proc. Entomol. Soc. Wash. 110: 130-143). [L. L. Deitz]

Explore the Treehoppers Taxon Pages
2011-08-19
Users may now Search or Browse our Taxon Pages, which include content for all valid treehopper genera and higher taxa. For all taxonomic names now entered in our database, the Search option returns either a taxon page (where available) or a report with nomenclatural details. Taxon pages and nomenclatural data for the species group, now in preparation, will eventually be accessible under the valid genus (Browse option) or directly through the Search option. Images are still needed to illustrate 16 genera: Acanthicoides, Aleptocentrus, Ananthasubramanianum, Aspasiana, Centrobelus, Demanga, Gargarina, Gonoconophora, Insitor, Insitoroides, Matonotus, Megaloschema, Neocentrus, Neomachaerotypus, Occator, and Spathenotus. We thank our Treehoppers' Nomenclatural Database Team (especially Matt Yoder and Katja Seltmann), our Taxa Content Team, and all who contributed images, acoustic calls, and other assistance. [L.L. Deitz and M.S. Wallace, 19 August 2011]

New Analysis of Smiliini and Telamonini by M. S. Wallace
2011-10-03
Wallace, M. S. 2011. Morphology-based phylogenetic analysis of the treehopper tribe Smiliini (Hemiptera: Membracidae: Smiliinae), with reinstatement of the tribe Telamonini. Zootaxa 3047: 1–42.) [available online]

Forthcoming Paper
2011-10-25
An early view of the following work was published online on 14 Oct 2011.

Yoshizawa, K. 2011. The treehopper's helmet is not homologous with wings (Hemiptera: Membracidae). Systematic Entomology [~5 pages]

Treehopper Specimens and Sculptures at the Montreal Insectarium
2011-11-29
Stephane Le Tirant, curator of the Montreal Insectarium, has a passion for treehoppers and provided a link to a list of the Insectarium's holdings of Membracidae. The Insectarium was recently reopened with a new permanent exhibit, including larger than life treehopper sculptures.


Robert Oelman's Extraordinary Images
2011-12-14
We are most grateful to nature photographer Robert Oelman for permission to include many of his amazing images of Neotropical treehoppers on our site. Robert's photos of treehoppers and other natural wonders may also be viewed at http://naturephotography.robertoelman.com/ (see the galleries: "Peruvian Amazon Insects & More" and "Treehoppers & More"); http://robertoelman.com/#/insects (see Hemiptera); and http://www.flickr.com/photos/roboelman/sets/72157624416166150/.

© Copyright 2011, Robert Oelman.

Latest on the Anatomy of the Treehoppper Pronotum
2012-01-09
A new article by Miko et al. (2012) provides further morphological evidence rebutting the work of Prud'homme et al. (2011), who suggested that the "pronotal" modifications of treehoppers may arise from fused appendages of the prothorax, serially homologous with the wings of the meso- and metathorax. The new paper, which includes an interactive image depicting the anatomy of the treehopper pronotum, concurs with Yoshizawa (2011) in concluding that the prothoracic superstructure in treehoppers does not correspond to a pair of fused wings, but represents the entire prothorax, except the forelegs.

Preview "Aliens of the Amazon: Treehoppers"
2012-01-15
A trailer for the film Aliens of the Amazon: Treehoppers may now be downloaded on online. This award-winning documentary was featured at the 2010 Treehopper Gathering in Little Orleans, Maryland, as noted in our news item on the event.

Congratulations to Dra. Evangelista
2012-03-13
We are delighted to report that Olivia Evangelista successfully defended her doctoral thesis "Systematics and Phylogenetic Analysis of the Treehopper Subfamily Heteronotinae Goding, 1926 (Hemiptera, Auchenorrhyncha, Membracidae)" on 23 February 2012. Working under the direction of Albino M. Sakakibara (Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil), Olive investigated phylogenetic relationships within Heteronotinae based on a combination of morphological and molecular data. The latter included six nuclear and mitochondrial markers (28S, H2A, H3, Wg, COI, ND1), which she compiled in Jason Cryan's lab at the New York State Museum, Albany. A major contributor to the Treehoppers website, Olive provided most of our content on the subfamily Heteronotinae. We are thrilled that Dr. Evangelista is part of our Team.

20th Annual Treehopper Gathering: 1-3 June 2012, Little Orleans, Maryland
2012-04-03
Join the 20th Annual Treehopper Gathering, 1-3 June 2012 (arrive Friday, depart Sunday), Little Orleans Campground, Little Orleans, Maryland. This casual event offers an opportunity to chat and collect with others passionate about treehoppers and related insects.

To register or obtain further information, contact Charles Bartlett. Campsites are somewhat limited, so please register with Charles if you wish to camp with our group. Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening.

Zoom in on Germar's Collection
2012-04-04
Thanks to the curators of the Benedict Dybowski Zoological Museum (Lviv National University, Ukraine), zoomable digital images of the Ernst Friedrich Germar Collection are now available online. For a number of years, this historically significant collection was thought to have been destroyed in World War II.

Celebrating Treehoppers for 20 Years
2012-06-11
The Annual Treehopper Gathering celebrated its 20th anniversary in Little Orleans, Maryland, on 2 June 2012. Charles Bartlett organized this year's event, which included 25 participants (Figs. 1 & 4). Four were among the original nine participants at the first gathering in Delaware (Fig. 2). Kelley Tilmon created a limited edition T-shirt just for the anniversary (Fig. 3), and Anthony Gonzon provided his delicious deep-fried turkey.



Fig. 1, 20th Gathering (left to right): back row, Anthony Gonzon, Ashley Kennedy, Cyrus Kasaaian, Meghan Irvine, Thomas Rounsville, Keith Bayless, Dennis Kopp, Adam Bell, Dawn Flynn, Lawrence Barringer, Lewis Deitz, Matt Wallace, Thomas R. Huff; middle row, Claire Ciafré, Katie Weglarz, D'Orsay Mancuso, Kelley Tilmon, Katja Seltmann, Mark Rothschild, Charles Bartlett, Brendan Morris, Julie Urban; front, Nikki January, Jason Cryan [see also Yon Visell, below, 4]. Fig. 2, First Gathering, Tom Wood's home, near the University of Delaware, 14 May 1993: top row, (left to right): Tom Greene, Tom Wood, Rex Cocroft, Stuart McKamey, Chris Dietrich; bottom row, Lewis Deitz, Kelley Tilmon, Jason Cryan, Charles Bartlett, Dianna McPherson [Bartlett].



Fig. 3, T-shirt design by Kelley Tilmon (with help from William Weeks Fowler). Fig. 4, Katja Seltmann and Yon Visell (photographer for Fig. 1).

Among the prize specimens collected at Little Orleans this year are adults and nymphs of Telamona ampelopsidis and a nymph of T. collina. Items of note from attendees include two treehopper checklists in press (Panama checklist by Dawn Flynn; Nearctic list by Lew Deitz & Matt Wallace) and several recent appointments: North Carolina Museum of Natural Sciences, Raleigh (Jason Cryan, Deputy Museum Director of Research & Collections, and Julie Urban, Assistant Director, Genomics & Microbiology Laboratory, Nature Research Center); Tri-Trophic Databasing Project, American Museum of Natural History (Katja Seltman, Project Manager [Charles Bartlett is a Co-PI on this NSF-funded project); University of Illinois (Brendan Morris, new M.S. student to study Caribbean treehoppers with Chris Dietrich); and National Museum of Natural History, Washington, D.C. (Dennis Kopp, new volunteer curator in Hemiptera section [recently retired from USDA/NIFA]).

Treehoppers featured on ID Source
2012-07-17
The Treehoppers website is currently featured on ID Source, a searchable database for locating online resources useful in identifying plant pests, diseases, and weeds. ID Source is published by Colorado State University as part of a cooperative agreement with the Center for Plant Health, Science and Technology (USDA APHIS PPQ). By helping those involved in plant protection find needed identification tools, ID Source supports the protection of U.S. agriculture and natural resources from the entry and spread of invasive plant pests.

Maruyama's 2011 book "Treehoppers: Incredible Insects"
2012-08-03
Maruyama, M. 2011. Treehoppers: Incredible Insects. Nature Editors, Gentosha, Japan. 80 pp. ISBN 978-4-344-02011-5. [in Japanese; figures with Latin scientific names including genus-species, tribe, and subfamily]

This attractive, full color book is an artistic celebration of the family Membracidae. Its astonishing images document nearly 100 New and Old World genera, representing 34 tribes, and serve as a model for high quality treehopper photography.

New Treehopper Checklists for Panama and the Nearctic Region
2012-08-16
New lists of the treehoppers of the Panama (Flynn 2012) and the Nearctic region (Deitz and Wallace 2012) show an overlap of fewer than 30 species, highlighting the distinctness of the Neotropical and Nearctic faunas.

Flynn, D. J. 2012. Checklist of treehoppers of Panama (Hemiptera: Membracidae) with a list of checklists and keys to the Nearctic and Neotropical fauna. Zootaxa 3405:35-63.

This checklist, the first for Panama, lists a total of 97 treehopper genera, 269 species, and 2 subspecies, including Flynn's new records. It also provides guides to species checklists and keys for various geographic areas within the Neotropical and Nearctic regions.

Deitz, L. L.; Wallace, M. S. 2012. Richness of the Nearctic treehopper fauna (Hemiptera: Aetalionidae and Membracidae. Zootaxa 3423: 1-26.

This overview of the taxonomic and regional diversity of the Nearctic treehopper fauna gives the Nearctic distribution for 280 species recorded from the region and documents the species richness of each Nearctic subregion, state, and province. One tribe, 16 genera, and 195 species are listed as Nearctic endemics.

Refinements to Taxon Searches
2012-08-20
We happily report that recent fine-tuning should vastly expedite navigating and searching our Taxon Pages. Enjoy the improvements.

Lewis Bartholomew Woodruff
2012-09-14
Lewis Bartholomew Woodruff, who described more than 15 treehopper species (mostly in the genera Cyrtolobus and Ophiderma), is featured in a recent note from the Tri-Trophic Thematic Collections Network. Yale University Library's Manuscripts and Archives includes a Guide to the Woodruff Family Papers. Z.P. Metcalf's 1944 Bibliography of the Homoptera (Auchenorhyncha) incorrectly attributed Woodruff's five publications on membracids to "Lois Bartolomew Woodruff."

Patience Please
2012-11-05
The Treehoppers website may be down for a time in the next few weeks due to essential work on the mx system. Know that folks are doing their best to complete the work before the end of November and to make this process as seamless as possible. Meanwhile, we appreciate your patience should things get a bit bumpy. [L.L. Deitz and M.S. Wallace]

T. K. Wood’s Treehopper Collection at Cornell University
2012-11-16
A news item from the Cornell University Insect Collection reports that the Thomas K. Wood Collection is now fully integrated into the University Insect Collection, bringing Cornell’s overall collection of Membracidae to 68 museum drawers. A complete listing of Cornell’s 497 membracid species is available online.

New Nearctic Records of Gargara genistae
2012-11-30
Wheeler, A. G., Jr., and E. R. Hoebeke. 2012. Gargara genistae (F.) (Membracidae) and Dictyonota fuliginosa Costa (Tingidae): new records of two immigrant, Scotch broom-feeding hemipterans in western North America. Proceedings of the Entomological Society of Washington 114(1):152-158.

This work includes new records of G. genistae from Canada (British Columbia: Vancouver Island) and the US: (Oregon : 5 counties; Washington: 6 counties). It also indicates that Metcalf and Wade's 1965 catalog record of the species for "Maine" (citing Poisson and Monguillon, 1931) refers to Le Maine, a former province of France. Thus, Deitz and Wallace's (2012) Nearctic records for G. genistae should be updated to: "Introduced: USA: Central & Eastern States: CT, NJ; Northwestern States: OR, WA. CANADA: Western Provinces: BC; [? Central & Eastern Provinces: ON]."

Treehopper Videos
2012-12-13
Several fascinating videos of treehoppers are available online, especially on YouTube. Please enjoy some of our favorites, but be advised that the identifications and commentaries have not been verified for scientific accuracy.

Treehopper Eggs Hatching

Publilia Molting to Adult (Stefano Di Criscio)

Treehopper Metamorphosis

Bocydium Signaling

Ennya Signaling

Treehoppers

Treehoppers of Costa Rica

Membracidae (INBio Costa Rica)

Umbonia Adult and Nymphs

Are Treehoppers Neotenous Leafhoppers?
2013-01-08
See Andy Hamilton's recent, thought-provoking article entitled: "Are Treehoppers Neotenous Leafhoppers?" (2012, American Entomologist, 58: 224-232).

New Interpretation of Paraneopteran Wing Venation
2013-01-10
For a new interpretation of the wing venation of treehoppers and related insects, see: Nel, A.; Prokop, J.; Nel, P.; Grandcolas, P.; Huang, D.-Y.; Roques, R.; Guilbert, E.; Dostál, O.; Szwedo, J. 2012a. Traits and evolution of wing venation pattern in paraneopteran insects. Journal of Morphology 273: 480-506.

New Work on the Tribe Talipedini
2013-02-01
Sakakibara, A. M. 2012. Taxonomic reassessment of the treehopper tribe Talipedini with nomenclatural changes and descriptions of new taxa (Hemiptera: Membracidae: Membracinae). Zoologia (Curitiba) 29 (6): 563–576. [available online]

Sakakibara notes a new unique feature of the Talipedini: the presence of latero-apical carinae on the posterior pronotal process. As redefined, this tribe includes 3 genera: Erechtia Walker, 1858 (Trinarea Goding, 1926, becomes a junior synonym), with 6 species (3 new); Pseuderechtia, new genus, with 1 species; and Talipes Deitz, 1975 (reinstated as valid), with 2 species. Other notable nomenclatural changes include the reinstatement of the genus Tropidoscyta Stål, 1869 (tribe Membracini) and the placement of 24 species previously included in Erechtia as of uncertain position within Membracini.

New Data on Jumping in Treehoppers
2013-02-18
Burrows, M. 2013a. Jumping mechanisms of treehopper insects (Hemiptera, Auchenorrhyncha, Membracidae). Journal of Experimental Biology 216:788-799. [online abstract; pdf (by subscription)]. Based on an analysis of high speed images of eight membracid species, the author concludes that jumping must involve "a power amplification mechanism in a catapult-like action."

Linnean Insect Collection Online
2013-02-22
Thanks to efforts of the Linnean Society of London, one can now examine the historically important collections of Carl Linnaeus online. His treehopper specimens are placed under the genus Cicada.

Of six treehopper species described by Linnaeus (see below), Cicada cornuta and C. foliata are represented in the collection, which also includes a mixed series of treehopper specimens identified as "Cicada carinata, a species described by Forster 1771a: 67 [now Entylia carinata (Forster)]."

Cicada cornuta Linnaeus 1758a: 435 [now Centrotus cornutus (Linneaus)]
Cicada crux Linnaeus 1758a: 435 [now Hemikyptha crux (Linneaus]
Cicada foliata Linnaeus 1758a: 435 [now Membracis foliata (Linnaeus)]
Cicada fronditia Linnaeus 1758a: 435 [now Stegaspis fronditia (Linnaeus)]
Cicada reticulata Linnaeus 1758a: 436 [now Aetalion reticulatum (Linnaeus)
Cicada squamigera Linnaeus 1758a: 435 [now Enchenopa squamigera (Linnaeus)]

Other Linnean species that resemble treehoppers are: Cicada aurita Linneaus, now placed in the family Cicadellidae [Ledra aurita (Linneaus)] and Cicada rhombea Linnaeus, now placed in the orthopteran family Tetrigidae [Phyllotetttix rhombeus (Linneaus)].

[L. L. Deitz, 22 February 2013]

Treehopper Vibes: David Rothenberg's Bug Music
2013-03-06
Treehopper vibes are featured in David Rothenberg's new book "Bug Music: How Insects Gave Us Rhythm and Noise" and album "Bug Music: the CD." Rothenberg is a professor of philosophy and music at the New Jersey Institute of Technology in Newark, New Jersey.

Eye-Popping Treehopper Design
2013-03-08
© Copyright 2013, Katja Seltmann.

Katja Seltmann, multi-talented member of our TREEHOPPERS team, created this stunning treehopper-inspired design for the cover of the 2013 Entomological Collections Network program. She is currently one of the officers and organizers of the ECN. We are most grateful to Katja for sharing her wonderful artwork, which is based on the genus Cladonota. Where and when can we order the wall paper, posters, and gift-wrap paper?

Join the 2013 Treehopper Gathering, Little Orleans, Maryland
2013-04-03
The 21st Annual Treehopper Gathering is scheduled for 31 May to 2 June 2013 (arrive Friday, depart Sunday), Little Orleans Campground, Little Orleans, Maryland. This casual event offers an opportunity to chat and collect with others passionate about treehoppers and related insects.

To register or obtain further information, contact Charles Bartlett. Campsites are somewhat limited, so please register with Charles if you wish to camp with our group. Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening.

Dr. Evangelista's Research on Heteronotinae
2013-04-22
Brazilian treehopper specialist Dr. Olivia Evangelista recently accepted a postdoctoral appointment at the Museu de Zoologia da Universidade de São Paulo, where she is continuing her research on the treehopper subfamily Heteronotinae. Currently, she is compiling needed DNA data in the molecular lab of Dr. Julie Urban (Nature Research Center, North Carolina Museum of Natural Science, Raleigh). While in Raleigh, Olivia presented two fantastic talks on treehoppers, including one (hosted by Science Comedian Brian Malow) inside the dome of the Center's Daily Planet. Images © Copyright 2013, Julie M. Urban, except 2nd and 4th of the smaller images © Copyright 2013, Jason R. Cryan.



Treehipster Tags and Other Creations of Brendan Morris
2013-06-18
One might say that Brendan Morris eats, breathes, and dreams treehoppers. A graduate student studying Caribbean treehoppers at the University of Illinois, Brendan uses spray paint, snow, and even French fries to create his treehopper art. Here are a few samples of his imaginative works, including a couple that have since melted or undergone digestion (images © Copyright 2013, Brendan O. Morris).



21st Annual Treehopper Gathering, Little Orleans, Maryland
2013-06-18
Treehopper enthusiasts and other entomologists came from as far away as South Dakota, Florida, Connecticut, and Kansas to attend the 21st Annual Treehopper Gathering (31 May to 2 June 2013) at Little Orleans, Maryland. As in past years, the camaraderie and food were great, but the weather this year was exceptionally fine, leading many to the camp's swimming pools.



Participants (above, from left to right) included Andrew Short, Matt Wallace, Dawn Flynn, Rachel Jansen, Jennifer Pohler, Lawrence Barringer, Bob Blinn, Cera Fisher, Yon Visell, Katja Seltmann, Lewis Deitz, Kelly Tilmon, Jason Owens, Dennis Kopp, Stuart McKamey, Mike Hennessey, Mark Rothschild, Anthony Deczynski, Drew Shaeffer, Amy Nazdrowicz, Nate Nazdrowicz, Shawn Dash, Ashley Kennedy, Charles Bartlett, and Anthony Gonzon. Photo © Copyright 2013, by Jason Owens.



Like many attendees, Charles Bartlett (organizer of the event; to the left in the above shots) and Anthony Gonzon (our head chief) took some time to go collecting. Photos © Copyright 2013, by Ashley Kennedy.

Societies of Sound in the Amazon
2013-08-29
A recent National Public Radio broadcast entitled Societies of Sound in the Amazon (Alex Chadwick, 19 August 2013, The Story) features Rex Cocroft's research on treehopper communication in the Amazonian rain forest of eastern Ecuador.

New Publication on the Treehopper Genera Antillotolania and Deiroderes
2013-08-30
McKamey, S. H.; Brodbeck, B. V. 2013a. Immature stages and hosts of two plesiomorphic Antillean genera of Membracidae (Hemiptera) and a new species of Antillotolania from Puerto Rico. ZooKeys 301: 1–12.

This online publication presents the first descriptions of the nymphs of Antillotolania and Dieroderes, along with the description of A. myricae, n. sp., from Myrica splendens in Puerto Rico. A table for identifying the four known species of Antillotolania is included.

New Findings on Speciation in Membracidae
2013-09-26
Stearns, F. W.; Tilmon, K. J.; Wood, T. K. 2013a. Felsenstein's "one-allele model" of speciation: The role of philopatry in the initial stages of host plant mediated reproductive isolation in Enchenopa binotata. Current Zoology 59 (5): 658–666.

Based on an 8-year experiment at the University of Delaware, this work explores the role of philopatry (a tendency to remain at one's birth site) as related to speciation in the Enchenopa binotata complex (Hemiptera: Membracidae).

The Neotropical Genus Lycoderides, stat. nov.
2013-10-23
Sakakibara, A. M. 2013. The genus Lycoderides Sakakibara, stat. nov., its composition and descriptions of new species (Hemiptera, Membracidae, Stegaspidinae). Revista Brasileira de Entomologia 57: 29-270. [available online]

Lycoderides (proposed as a subgenus in 1972) is elevated to the level of genus. A key and illustrated descriptions are given to identify the 24 included species, 6 of which are new. The work also includes new geographic records for some species of this Neotropical genus.

Amazing Treehoppers of Ecuador (18 to 22 November 2013)
2013-11-15
Dear exotic insects enthusiasts! You haven't seen it all! Last month, we were fortunate to stumble across an undiscovered world, as far as insects are concerned, not far from the Napo River in tropical Ecuador, and I have returned to tell, or rather to show, the tale. The best of my photos of these amazing creatures will be presented on my Flickr website starting next Monday, the 18th of November, and will continue on weekdays for four weeks. There will be between 4 and 8 photos posted each day. Treehoppers will occupy the first week.

Don't miss this show by some of the most wonderful and "self-made artists" of the natural world! -- Robert Oelman, 14 November 2013

First Description of Immature Alcmeone
2014-01-09

Alcmeone robutus (Butler), last instar nymph. © 2013, by Federico Leniconi-Neto

Lencioni-Neto, F.; Sakakibara, A. M. 2013a. Alcmeone robustus (Hemiptera: Membracidae: Darninae): description of the last-instar nymph and biological notes. Zoologia (Curitiba) 30(5): 471-474. [available online] This work describes the fifth instar nymph of A. robustus and provides notes on the biology and behavior of the species as observed on Pera sp. in Brazil.

Treehoppers and Pop Culture
2014-01-15
The treehopper genus Bocydium was recently listed among five "theatrical" animals that look like Lady Gaga, and it seems that more than one treehopper has been compared to the stylish recording icon.

But, treehopper fashion is not exclusively for Lady Gaga. Anyone can now purchase treehopper underwear as well as other "popular" treehopper products.

Treehoppers have also found a place in popular films. Treehoppers are mentioned in "Master and Commander: The Far Side of the World," a historical drama co-written and directed by Peter Weir, and treehopper sounds were used in " "A.I. Artifical Intelligence," a science fiction film directed by Steven Spielberg.

Last, but not least, we recently discovered an wonderful work of art entitled the "Green Treehopper," and hope someone can tell us the name of the artist who created it.

Species Taxon Pages for Monobelini and Nessorhinini
2014-01-28
Populating the Treehoppers Database is a slow process, and we greatly appreciate your patience as we develop this resource. Treehopper Taxon Pages are available for all genera and higher groups, and we have begun entering Species Pages for selected taxa.

Thanks to the assistance of Brendan Morris, our taxon searches now provide Taxon Pages for all valid species in the Caribbean tribes Monobelini and Nessorhinini. These pages may include synonyms (generated from our nomenclatural data), selected references (these will eventually link to a full citation), images, distribution data, diagnosis, and so forth.

Note that Nomenclatural Reports are displayed for invalid taxa (genus group or higher) and for those species (valid or invalid) for which only nomenclatural data have been entered. The second line of text will state: "This is a nomenclatural report, not a taxon page." Olivia Evangelista has prepared nomenclatural reports for all species of the Neotropical subfamily Heteronotinae.

[L. L. Deitz, O. Evangelista, S. H. McKamey, M. J. Rothschild, and M. S. Wallace]

Membracid Types at North Carolina State University
2014-02-03
Bob Blinn, collection manager of the NCSU Insect Museum, is developing an online list of NCSU's types. The collection holds type material of 44 treehopper species, including 27 holotypes. For primary types, the listing includes the original binomen, author(s), and literature citation, along with information quoted from the data labels. For some species, images of the type and its labels are available online, as shown below for the holotype of Aconophora elongata (photos Copyright © 2013, by Robert L. Blinn).



Aconophora elongata Dietrich In Dietrich & Deitz 1991. N. C. Agric. Res. Ser. Tech. Bull. 293:112.
Holotype: "PERU/Chancha/mayo; 21-vii-60/J. Salazar; on/Leguminosae; DIETRICH/RESEARCH/4-84-373c; HOLOTYPE/Aconophora/elongata/Dietrich"
Paratypes, 8: Peru

Green Treehopper: Artist Revealed
2014-02-03

Copyright © 2013, by Michihiro Matsuoka

The creator of the amazing Green Treehopper is Japanese sculptor Michihiro Matsuoka. This 200x200x60 mm work incorporates "resin clay, aluminum rod, acrylic, and plastic plate." Inspired by childhood memories of broken devices, Matsuoka's fantastic creations are part animal and part machine. His work has been featured in exhibitions from Japan to San Diego, New York, Germany, and Belgium.

We are grateful to Michihiro and his agent, Ben Wouters, for permission to use the photo (above) and to another artist, Dan Beaulieu, for connecting the sculptor to the sculpture (see prior news item: "Treehoppers and Pop Culture").

Treehoppers on Stamps
2014-02-25
Leafhopper specialist Paul Freytag kindly provided the following data related to three known postage stamps that depict treehoppers. A zealous collector of Auchenorrhyncha on stamps, Paul is an emeritus professor at the University of Kentucky, Lexington. As indicated on the images, the stamps are in the collections of John R. Meyer and Paul H. Freytag.

Family Membracidae

1978 Ivory Coast: issued 26 August 1978: Centrotus cornutus (Linnaeus). 60 fr., Scott #478. Also known as imperforate & deluxe sheet.

1998 Tanzania: issued 27 November 1998: Umbonia sp., labeled thornbug on stamp. 1500 sh., Scott #1748.

1999 Republic of Guinea: issued 11 November 1999: Umbonia crassicornis (Amyot and Serville), male, not labeled on stamp. 450 fr., Scott #1563.



Observing Treehoppers in Chile, Argentina, and Brazil
2014-02-27
Chris Dietrich recently returned from a 7-week trip to South America (Brazil, Chile, and Argentina) where he collected and photographed many interesting treehoppers, including members of the family Melizoderidae, restricted to Chile and Argentina, and the strange nicomiine genus Holdgatiella, a Chile endemic. Other highlights of the trip were visits to national parks in and around Rio de Janeiro that preserve remnants of the Brazilian Atlantic Forest, home to many interesting treehoppers, including the heteronotine genus Iria.


Nymphs and adults of Melizoderes spp. (first row: left, nymph; right, adult; second row: left, adult), Llanquihea pilosa Linnavuori and DeLong (second row: right, female; third row: left, nymphal female; right, male), and Holdgatiella chepuensis Evans (fourth row: right, adult; left, nymph), all photographed in Chile - © Copyright 2014, by Christopher H. Dietrich.

Treehopper News of 1 April 2014
2014-04-01
What time is it? It's Treehopper Time!

© Copyright 2014, Stuart H. McKamey.

Was the Secretary of Defense trying to identify treehoppers?… as we know, there are known knowns; there are things we know that we know. There are known unknowns; that is to say, there are things that we now know we don't know. But there are also unknown unknowns – there are things we do not know we don't know.” Donald Rumsfeld, February 2002

What Charles Darwin would have said on this first day of April:Had the Treehoppers Website been available when I was alive, I would have used it every day.

Register for the 2014 Treehopper Gathering, Little Orleans, Maryland
2014-04-04
The 22nd Annual Treehopper Gathering is scheduled for 27 to 29 June 2014 (arrive Friday, depart Sunday), Little Orleans Campground, Little Orleans, Maryland. This casual event offers an exceptional opportunity to meet and collect with others passionate about treehoppers and related insects.

To reserve a campsite with our group or to obtain further information, contact Charles Bartlett. Advanced registration with Charles is essential to assure that you have a campsite.

Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening. Bring your cameras, macro lens, collecting vials, unidentified specimens, and bathing suits!

Revised Link to the Germar Collection
2014-04-30
We here provide an updated link to digital images of the Ernst Friedrich Germar Collection. For a number of years, this historically important collection was thought to have been destroyed in World War II. As we reported in March 2012, zoomable images of the collection are available online thanks to the curators of the Benedict Dybowski Zoological Museum (Lviv National University, Ukraine).

Major Revision of the Genus Enchenopa
2014-05-07
Strümpel, H.; Strümpel, R. 2014a. Revision der amerikanischen Membracidengattung Enchenopa (Hemiptera: Auchenorrhyncha: Cicadomorpha: Membracidae) mit Beschreibungen neuer Arten. [= Revision of the American treehopper genus Enchenopa (Hemiptera: Auchenorrhyncha: Cicadomorpha: Membracidae) with description of new species.] Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 17(191): 1-137. [in German with English abstract]

The genus Enchenopa is redefined to include 51 valid species (21 new), with Campylenchia treated as a junior synonym. Nomenclatural changes at the species level include seven new synonymies and the reinstatement of four species as valid. Nine lectotypes are designated. The work includes a key to the species and a total of 102 figures (most depicting more one than structure).

Inferring Treehopper Phylogeny from Genetic Barcode Data
2014-06-05
Hamilton, K. G. A. 2014a. How quickly do insects evolve? American Entomologist 60(1): 14-22.

This paper presents a phylogeny based on genetic barcode data for “25 species groups representing 36 telamonine treehoppers (Membracidae).” Based on a neighbor-joining analysis of a 658 base-pair segment of the COI gene, the results support the view that treehopper genera defined solely on the basis of pronotal similarity may not be monophyletic. Genetic barcoding is indeed “still in its infancy” (as noted in the paper), so stay tuned.

Two New Umbelligerus from Panama
2014-06-05
Flynn, D. J. 2014a. Review of the genus Umbelligerus Deitz with descriptions of two new species from Panama and key to adults (Hemiptera: Membracidae). Proceedings of the Entomological Society of Washington 116(2): 145-154.

This work describes Umbelligerus stockwelli<> and <>U. convergens, n. spp. With a total of five Neotropical species, Umbelligerus is reported from Brazil, Costa Rica, Panama, Peru, and Venezuela.

27- 29 June 2014: Treehopper Gathering: Little Orleans, Maryland
2014-06-05
This is a reminder that the 22nd Annual Treehopper Gathering is scheduled for 27 to 29 June 2014 (arrive Friday, depart Sunday), Little Orleans Campground, Little Orleans, Maryland. This event offers an exceptional opportunity to meet and collect with others passionate about treehoppers and related insects. To reserve a campsite with our group or to obtain further information, contact Charles Bartlett. Advanced registration with Charles is essential to assure that you have a campsite. Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening. Bring your cameras, macro lens, collecting vials, unidentified specimens, and bathing suits!

New World Treehopper Introduced to Control Lantana in Australia Damages Fiddlewood Trees
2014-06-05
Queensland Department of Agriculture, Fisheries and Forestry. 2013a. Aconophora on fiddlewood trees: Aconophora compressa. Department of Agriculture, Fisheries and Forestry (Queensland, [Australia]), Biosecurity Queensland Fact Sheet, Pest Animal PA27: [1-3]. [available online]

Introduced from Mexico in 1995 to control lantana in Queensland and New South Wales, Aconophora compressa may cause die-back and even defoliate fiddlewood trees.

2014 Treehopper Gathering: Little Orleans, Maryland
2014-07-14
The 2014 Treehopper Gathering (27 to 29 June) attracted entomologists from Brazil, Ecuador, and as far as Florida, Kansas, New York, and Connecticut in the United States. See group photo, below, left to right (b= back; f= front): Charles Bartlett, bAnthony Deczynski, fAlexis Park, bAndrew Short, Yuri Park with son fDuncan, bAdam Bell, Robert Clark, bKeith Bayles, fLuis Camacho, bLewis Deitz, Stuart McKamey, bMike Hennessey, Olivia Evanelista, fLogan Fish, Matt Wallace, fCamille McKamey, bCorey Janusz, Cera Fisher, bGlenn E. Miller, Gretha dos Santos, bStephen Kloiber, Dawn Flynn, bMitchell Porter, Mark Rothschild, bMichael (Drew) Sheaffer, Nate Nazdrowicz, bAnthony Gonzon [Lawrence Barringer, not shown]. Smaller images show selected shots from the Little Orleans area. Images © Copyright 2014, by Robert E. Clark.





Attendees were among the first to see printed copies of "A review of the Planthoppers (Hemiptera: Fulgoroidea) of the United States,” by Bartlett, O’Brien, and Wilson. "Hot off the press," this impressive and richly illustrated volume includes: a checklist of the 12 families, 167 genera, and 914 species occurring in the contiguous United States, plus 5 further genera (26 species) occurring in Canada and Alaska; keys to the families and genera; summaries of distribution; and an update to Metcalf’s bibliography. Another hot topic of discussion was the status of the genus Campylenchia, recently placed as a junior synonym of Enchenopa (Strümpel and Strümpel 2014a), but supported as a distinct genus in a prior molecular study (Lin, Danforth, and Wood 2004a). Attending systematists noted the need for sound phylogenetic analyses, as well as an examination of evidence from studies of life history, behavior, immatures, male mating calls, and host plant associations.

Participants collected a variety of insects at lights in the evenings. Images © Copyright 2014, by Luis Camacho.


This year’s gathering was purposely timed in order to sample species that occur a little later in the collecting season. Thus some later season species were found including a few immatures of the tribe Telamonini. Recognizing the importance of features of immatures in treehopper systematics, Stu McKamey is preparing an identification key to the nymphs of the New World tribes and genera, while Matt Wallace is focusing on the nymphs of the Telamonini. As it stands, the 2015 Gathering will likely be scheduled for early June. We hope to announce the final dates 2 or 3 months before the event.

Dysnycritus and the new genus Allodrillus (Membracidae: Heteronotinae)
2014-08-15
Evangelista, O.; Flórez-V, C.; Sakakibara, A. M. 2014a. The identity of the treehopper genus Dysnycritus Fowler, with descriptions of new related taxa (Homoptera: Membracidae: Heteronotinae). Zootaxa 3847(4): 495-532.

[Dysnycritus is redefined to include only D. intectus Fowler. Allodrilus, n. gen., includes 8 spp. (7 new). Nomenclatural changes involve species moved from Dysnycritus to Smiliorachis (resulting in 2 new combinations and a new synonymy) and to Allodrilus (1 new combination). This richly illustrated work includes a key to the males of Allodrilus.]

Another Dimension to Treehopper Subsociality
2014-08-25
Camacho, L.; Keil, C.; Dangles, O. 2014a. Factors influencing egg parasitism in sub-social insects: insights from the treehopper Alchisme grossa (Hemiptera, Auchenorrhyncha, Membracidae). Ecological Entomology 39(1): 58–65.

This work explores the impact of maternal care on egg-parasitism in A. grossa. Females of this treehopper guard their egg masses, using their bodies and ponota to cover the eggs. Moreover, females actively ward off egg parasites by fanning their wings and kicking with their club-shaped hind legs. Camacho et al. found that increased numbers of females aggregated on a single host plant lower the risk and levels of egg parasitism through a “herd dilution effect.” The impact of possible cooperation among egg-guarding females adds another dimension to subsociality in the treehopper tribe Hoplophorionini.

Pyrgonota bifoliata: a Species Complex
2014-08-26
Su, Y.-C.; Wang, J.-F.; Villanueva, R. J. T.; Nuñeza, O. M.; Lin, C.-P. 2014a. Hopping out of Mindanao: Miocene-Pliocene geological processes and cross-island dispersal as major drivers of diversity for Philippine treehoppers. Journal of Biogeography 41(7): 1277-1290.

Work on the biogeographical history of Pyrgonota bifoliata suggests that Miocene-Pliocene geological events and related cross-island dispersal contributed to species diversity in the Philippines. Using phylogenetic reconstruction based on two genes (mitochondrial cytochrome c oxidase 1 subunit [cox1] and nuclear wingless [wg]), the authors found evidence of nine cryptic species that form the Pyrgonota bifoliata species complex.

Immatures and Host Plants of Telamonine Treehoppers
2014-10-24
Wallace, M. S. 2014a. The host plants of the Telamonini treehoppers (Hemiptera: Membracidae: Smiliinae) and the first diagnoses of nymphs for 14 species. Zootaxa 3878 (2): 146–166.

Owing to their cryptic shapes and coloration, nymphs of the treehopper tribe Telamonini (with 68 species) closely resemble parts of the plants on which they feed and develop. Wallace’s work provides the first comprehensive list of telamonine host plants along with useful morphological diagnoses and color photos for identifying the fifth instar nymphs of 15 species. The cryptic nymph of Archasia belfragei, below, is hidden in plain view (photo © 2013, by Matthew S. Wallace).



Members of this tribe are recorded from 80 host plant species in 41 genera and 22 families. The total number of hosts species/treehopper ranges from a single host (in 12 telamonine species) to 29 different hosts (in Telamona monticola). Indeed, many telamonines are recorded from more than one plant genus. The genus Quercus serves as a host for 45 of the tribe's 68 species.

Master of Science Thesis on the Membracidae of Colombia
2014-11-20
González Mozo, L. C. 2014. Sinopsis de Membracidae (Hemiptera: Membracoidea) de Colombia, relacionados con ecosistemas agrícolas. Maestria en Ciencias Agrarias--Entomología, Universidade Nacional de Colombia, Bogotá, Colombia. [iv] + viii + 145 pp. [M.S. thesis; available online]

Wild, Weird, and Wired
2014-12-10
Check out treehopper facts, images, videos, and sounds featured in Absurd Creature of the Week: This Is an Actual Insect. This Is Not a Joke, by Matt Simon (Wired.com: 5 December 2014).

Membracid Types at the Museu de Zoologia da Universidade de São Paulo
2014-12-15
Evangelista, O.; Santos, G. I. M. dos; Lamas, C. E. 2014a. An annotated catalogue of the Membracidae types in the Museu de Zoologia da Universidade de São Paulo, Brazil (Hemiptera: Auchenorrhyncha: Cicadomorpha). Zootaxa 3895(1): 1–30.

This useful catalogue documents 106 primary and 171 secondary membracid types held at the Museu de Zoologia da Universidade de São Paulo (MZSP). These specimens represent 82 nominal species in 17 treehopper tribes (8 subfamilies). The authors include notes on taxonomy, extensive literature sources, and for each type: precise label data and notes on the condition of the preserved material.

Colombian Membracidae: 474 Species in 116 Genera
2015-01-15
Flórez-V., C.; Wolff, M. I.; Cardona-Duque, J. 2015a. Contribution to the taxonomy of the family Membracidae Rafinesque (Hemiptera: Auchenorrhyncha) in Colombia. Zootaxa 3910 (1): 1–261. [in English with Spanish abstract]

With color images, diagnoses, and a key for identifying the membracid genera of Colombia, this publication increases the numbers of membracids reported for that country to 116 genera and 474 species. This research was based on the literature, field collections and observations (2011 to 2014), and the examination 3744 specimens held in ten entomological collections in Colombia. The authors provide data on biology and distribution for each genus and also highlight features differentiating the genera Anobilia and Stilbophora (Smiliinae: Tragopini), referring six species from the former to the latter as new combinations.

New Telamonine Treehopper from Costa Rica: Mutilifolia nishidai
2015-01-23
Wallace, M. S. 2015a. A striking new treehopper genus Mutilifolia (Hemiptera: Membracidae: Smiliinae: Telamonini), from Costa Rica. Zootaxa 3914 (1): 83–88.



© 2015, Kenji Nishida

The new genus and species Mutilifolia nishidai differs from those of all known telamonine genera in bearing two recurved teeth on the male style clasp. The species is named in honor of treehopper enthusiast Kenji Nishida, who collected the type material at the Monteverde Biological Station in the highlands of Costa Rica. Kenji regularly contributes exceptional photos of treehoppers to this Website, including the images above. The pronotum of Mutilifolia--folicaceous, highly elevated, largely green--is similar to that in species of Antianthe, Archasia, and Hemicardiacus.

Neotropical Treehoppers: Recent Discoveries
2015-02-09
Sakakibara, A. M. 2014. A new species of Oeda (Hemiptera: Membracidae: Stegaspidinae) from Madre de Dios, Peru. Zoologia 31 (6): 557–560. [available online]

This work describes a new Peruvian treehopper, Oeda (Oeda) mielkei.

Lencioni-Neto, F.; Sakakibara, A. M. 2014 . Registro da ocorrência de Enchenopa beebei (Haviland) (Hemiptera, Membracidae) para o Brasil. [Registration of occurrence of Enchenopa beebei (Haviland) for Brazil.] Revista Univap (São José dos Campos-SP-Brasil) 20 (35): 86-92. [available online]

Enchenopa beebei (Haviland, 1925), previously known from Guiana, is reported from remnants of the Atlantic Forest, State of São Paulo, Brazil.

Plea for Care in Pronouncing Treehopper Names
2015-03-31
It has come to our attention that the careless pronunciation of certain treehopper names may offend or confuse workers here and abroad. We are not talking about George Willis Kirkaldy’s generic names for many insects, such as “Ohchisme, Dolichisme, Elachisme, Florichisme, Marichisme, Nanichisme, Peggichisme, and Polychisme,” all of which are considered sweet because the suffix “-chisme” (Greek) is pronounced “kiss-me”--nor even his treehopper genus “Alchisme,” which was perhaps more provocative when first proposed in 1904. Rather we refer to the common American pronunciation of the genus “Heranice,” which, if carelessly pronounced, may sound just as offensive as the usual American pronunciation of the name of the seventh planet from the sun. Here, we endorse the Portuguese pronunciation, in which the initial “H” is silent and the final “ce” is pronounced like the English word “see.”

In contrast, Spanish speakers would do well to Anglicize the unfortunate genus name “Llanquihuea” (based on Llanquihue, a province in Chile), thus not pronouncing the double “L” as a “Y.” Apparently, in Spanish this name sounds either hilarious or vulgar, depending on one’s point of view.

To reduce the large number of useless petitions to the International Commission on Zoological Nomenclature requesting the suppression of these names, we urge workers to preserve the dignity of treehoppers by pronouncing their names with care, decency, and good taste.

--Treehopper Specialists’ Declaration of 1 April 2015

23rd Annual Treehopper Gathering: 5-7 June 2015, Little Orleans, Maryland
2015-04-07
Join the 2015 Treehopper Gathering, set for 5 to 7 June (arrive Friday, depart Sunday), at the Little Orleans Campground, Little Orleans, Maryland. This casual get-together provides a pleasant opportunity to meet, swap stories, and collect with others passionate about treehoppers and related insects.

To reserve a campsite with our group or to obtain further information, contact Charles Bartlett. Advanced registration with Charles is essential to assure that you have a campsite.

Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening. Bring your collecting vials, insect nets, unidentified specimens, cameras, macro lens, and bathing suits!

False Records of Telamona in South America
2015-04-15
McKamey, S. H.; Wallace, M. S. 2015a. The identity of three South American “telamonine” treehoppers (Hemiptera: Membracidae). Proceedings of the Entomological Society of Washington 117(1): 22-26.

Telamona ruificarinata Fowler, from Colombia, is referred to the genus Alchisme, with A. insolita Creão-Duarte and Sakakibara as a new junior synonym. Also, two species of Telamona described from Brazil–T. celsa Goding and T. turritella Buckton–are confirmed to be junior synonyms of the Nearctic species T. monticola (Fabricius) and Heliria praealta Fowler, respectively, their original type-locality data being incorrect.

Biodiversity Heritage Library Holds Broomfield’s 1971 & 1976 Works on Treehoppers
2015-04-20
Broomfield, P. S. 1971a. A catalogue of the membracid types (Homoptera: Membracidae) in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology 25(8):325-386. [available online]

Broomfiled, P. S. 1976a. A revision of the genus Amastris (Homoptera: Membracidae). Bulletin of the British Museum (Natural History), Entomology 33(4): 347-460. [available online]

Spiny Hopperling
2015-05-01
This spiny treehopper nymph is one of Katja Seltmann’s new series of bug inspired drawings. We hope it makes you smile.


Copyright © 2015, by Katja Seltmann.

Updates of Treehopper Photographers and Other Collaborators
2015-05-12
We gratefully acknowledge the many generous individuals who provide digital images of treehoppers or assist in identifying images for us. While most of these collaborators are already included in our Acknowledgments, we list below new supporters since our last update. The most recent image contributed is this stunning photo of a Membracis species [identification revised on 26 May 2015], taken in Brazil by Weiwei Zhang.


© 2015, by Weiwei Zhang

Collaborating photographers: Matthew A. Bertone (Department of Entomology, NCSU, Raleigh); Adam J. Bell (University at Albany, SUNY, Albany, New York); Robert E. Clark (Wesleyan University, Connecticut); Ashley M. Dennison (East Stroudsburg University, Pennsylvania); Olaf H. H. Mielke (Universidade Federal do Paraná, Curitiba-PR, Brazil); Dale Lee Denham-Logsdon (Dalesmicro, Toluca, State of Mexico, Mexico); Camilo Flórez-Valencia (Universidad de Antioquia, Colombia); Gustavo F. Morejón J. (Cuenca, Ecuador); Eli S. Wyman (American Museum of Natural History, New York); Weiwei Zhang (Chongqing, China). Other collaborators: Dawn J. Flynn (Schiele Museum of Natural History, Gastonia, North Carolina); Ai-Ping Liang (Key Laboratory of Zoological Systematics and Evolution, Chinese Academy of Sciences, Beijing).


Umbonia crassicornis (Amyot and Serville) © 2013, by Matthew A. Bertone

Reminder: June 2015 Treehopper Gathering in Little Orleans, Maryland
2015-05-12
The 2015 Treehopper Gathering will be 5 to 7 June (arrive Friday, depart Sunday), at the Little Orleans Campground, Little Orleans, Maryland. To reserve a campsite with our group or to obtain further information, contact Charles Bartlett. Advanced registration with Charles is essential to assure that you have a campsite. Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening.

2015 Treehopper Aggregation at Little Orleans, Maryland
2015-06-17
The 23rd Annual Treehopper Gathering (5 to 7 June, Little Orleans, Maryland) brought together treehopper enthusiasts from Delaware, Florida, Kansas, Maryland, New York, North Carolina, Pennsylvania, South Carolina, and South Dakota.

Fig. 1. Group photo of 2015 attendees., left to right (b= back; f = front): Charles Bartlett, Ruth Moscovitchf, Vinton Thompson, Mark Rothschild, Kelley Tilmon, Andrew Shortb, Dennis Kopp, Stuart McKameyf, Adam Bellb, Anthony Gonzon, Nate Nazdrowiczf, Lawrence Barringerb, Anthony Deczynski, Ashley Kennedyf, Yuri Park, Lewis Deitzb, Alexis Park, Duncan Parkf.


© Copyright 2015, by Nate Nazdrowicz.

The pleasant weather was slightly cooler than usual for early June, perfect for camping, socializing, and collecting in the woods of western Maryland. Our Saturday evening feast, always a highlight of the weekend, featured turkey (deep fried), steak, venison (all prepared by Anthony Gonzon), and teriyaki chicken (prepared by Yuri Park), as well as salads, various veggies (including baked potatoes and sweet potatoes), and desserts. Between meals, picnic tables were generally cluttered with drawers, boxes, and vials of treehoppers and other insects, including fresh insects being processed, hundreds of preserved specimens to be lent, returned, or donated to attendees or others at their home institutions, and an incredible display of treehoppers that Mark Rothschild collected this spring. (See photos 2-3).

While Vinton Thompson was off collecting spittlebugs, his wife Ruth Moscovitch sketched the valley view from Saint Patrick’s, a church surrounded by the Little Orleans Campground. Near this church, vines of Virginia creeper host a spectacular treehopper, Telamona ampelosidis. (See photos 4-6).

Figs. 2-7. 2, Adam Bell (left to right), Andrew Short, Charles Bartlett, and Lawrence Barringer collecting at the group campsite. 3, Charles Bartlett, Anthony Deczynski, and Lawrence Barringer under the picnic shelter. 4, Ruth Moscovitch with her sketch from Saint Patrick’s Church. 5. Saint Patrick’s Catholic Church and historic cemetery where a number of Irish workers involved in the construction of the C&O Canal are buried. 6, Telamona ampelosidis, a handsome treehopper that feeds on Virginia creeper, Parthenocissus quinguefolia (L.) Planchon. 7. Unexplained stick figures composed of small, dead wooden twigs tied together with waxed and unwaxed dental floss.




© Copyrights 2015, by Ashley Kennedy (photos 2 and 3), Lewis L. Deitz (photos 4, 5, and 7), and Mark J. Rothschild (photo 6).

Little Orleans is not far from Burkittsville, Maryland, where the terrifying movie “The Blair Witch Project” was allegedly filmed. This year, as in 2000 (following the film’s release in 1999), witchy stick figures mysteriously appeared around our campsite (Fig. 7), even in the absence of our beloved colleague and occasional mischief-maker Matt Wallace. Unlike the horror film, the Treehopper Gathering ended happily and without incident.

The 2016 Gathering will likely be scheduled for early June. We hope to announce the final dates 2 or 3 months before the event.

Recent Papers
2015-07-03
Harvey, A.; Wheeler, A. G., Jr. 2015a. Vanduzea segmentata (Fowler) (Hemiptera: Membracidae): seasonality and habits in the southeastern United States, with review of its U. S. distribution and host plants. Proceedings of the Entomological Society of Washington 117(2): 135-150.

The authors present a wealth of information on the life history and taxonomy of V. segmentata, including new host plant and state (Alabama and Georgia) records. Described from Neotropical Mexico (Guerrero, Tabasco, Veracruz), the species also occurs in Guatemala and Panama and was introduced to Hawaii (Harvey and Wheeler 2015a). Nearctic records now include: [? MEXICO (Nearctic): Nuevo León, Tamaulipas]; USA: Southwestern States: AZ, TX; Central & Eastern States: AL, FL, GA, LA (Deitz and Wallace 2012a; Harvey and Wheeler 2015a).

Hamilton, K. G. A. 2015a. Anatomy: the poor cousin of morphology. American Entomologist 61(2): 88-95.

This paper includes brief notes on Membracidae.

A Newly Described Pest of Soybeans in Brazil
2015-07-14
Two species of the genus Ceresa are associated with soybeans in Brazil: Ceresa brunnicornis (Germar 1835) from the southern region and Ceresa atlantica Andrade 2015, n. sp., from the east and northeast (Andrade 2015a). The latter species has been called the “soybean buffalo treehopper” in the applied entomological literature (Zanella 2007a). Andrade (2015a), who formally described this economically significant treehopper, kindly provided an original map (below), which summarizes the known distribution of C. atlantica in northeastern Brazil. [Map © 2015, by Gabriel Simões de Andrade]



Andrade, G. S. de. 2015a. A new species of Ceresa Amyot & Serville (Hemiptera: Membracidae) associated with soy culture in Brazil. Entomological News 125 (1): 47-51.

Zanella, M. 2007a. Búfalos na soja. Cultivar Grandes Culturas 97: 24-26.

Accessing the Archives of Early Literature on Treehoppers
2015-07-31
Z. P. Metcalf used unique bibliographic identifiers (author, year plus letter suffix) for works cited throughout his catalogues and bibliographies of the Auchenorrhyncha, the insect suborder of Hemiptera that includes treehoppers. Covering works published through 1955, his identifiers were widely used by many subsequent workers and continue to be used on such online resources as the Treehoppers Website and the Planthopper Digital Library.

DrMetcalf and the Internet Archive. Using the searchable DrMetcalf database, one can easily expand Metcalf’s identifiers to full bibliographic citations that provide essential data for locating and viewing the publications. With journal articles, the full journal title, volume, and start page are often key to locating the publication (individual article titles and authors may or may not be indexed in databases). Indeed, hundreds of early publications, including rare books and obscure serials, are now available online through the Biodiversity Heritage Library and other resources of the Internet Archive.

Index to Treehopper Publications in the Digital Library for Pre-1940 Planthopper Literature. Thanks to the efforts of Charles Bartlett, more than 140 references on treehoppers (listed below by Metcalf’s identifiers) are currently available through the Digital Library for Pre-1940 Planthopper Literature: Amyot and Serville 1843a; Ashmead 1888a; Atkinson 1885b, 1886b; Baker 1924b; Ballou 1936b; Barber 1914a; Berg 1879e, 1880a, 1881b, 1883c, 1884d; Bergroth 1915a; Britton 1920a, 1938c; Buckton 1890a, 1891d; Burmeister 1835a; Costa Lima 1932a, 1936a; Crawford 1914a; Crevecoeur 1905a; Dallas 1867a (1954 reprint), 1870a (1954 reprint); Distant 1888a, 1908g, 1912b, 1914i, 1914j, 1916a; Dohrn 1859a; Dozier 1928a, 1931a; Fabricius 1775a, 1798a; Fallén 1826a; Fieber 1872a; Germar 1833a; Glover 1877a; Haupt 1929b; Hayes 1922a; Heidemann and Osborn 1917a; Henshaw 1903b; Herrich-Schaefer 1935a; Jensen-Haarup 1920a; Johnson 1930a; Johnson and Fox 1892a; Kershaw 1913e; Kershaw and Muir 1922a; Kirkaldy 1900b, 1900c, 1901a, 1901d, 1901e, 1902a, 1903b, 1903c, 1903d, 1904c, 1906c, 1907d, 1908d,1909c; Knowlton 1929b; Kusnezov 1829b; Leonard 1933b; Lethierry 1881b, 1881c, 1881d, 1881e, 1890a; Licent 1912a; Linné 1758a (1894 reprint [= Linnaeus 1758a]); Matsumura 1905a, 1936a, 1938a; Mayr 1884a; Melichar 1896a, 1903b, 1905a; Metcalf 1915a; Motschulsky 1863a; Muir 1926a; Osborn 1898a, 1900f, 1904a, 1909a, 1921a; 1922c, 1926e, 1929b, 1935a; Osborn and Drake 1922a; Oshanin 1908b, 1912a, 1913b; Perkins 1907a; Popenoe 1885a; Provancher 1889a, 1890b; Sabransky 1912a; Sahlberg 1871a; Say 1830a; Schmidt 1911b, 1924b, 1927a, 1931b; Smith 1890a; Spinola 1852a; Stål 1853b, 1854b, 1855a, 1859b, 1862b, 1862e, 1863a, 1866a, 1869a, 1870c; Uhler 1872a, 1876a; Valdes Ragues 1910a; Van Duzee 1890a, 1892d, 1893b, 1894e, 1905a, 1906a, 1907a, 1908b, 1909a, 1909c, 1912a, 1912b, 1914a, 1916a, 1917a, 1917b, 1923a, 1925b: Walker 1851a, 1858a, 1858b.

Speaking of “archives,” to retrieve prior Treehopper News items, go to the bottom right of the current “News,” click on “archive of past news,” and use the “find” option (such as "control + f") appropriate to your computer to search for a keyword (subject, taxon or author names) in our archives. For example, this news item may be retrieved using any of the author names listed above or other keywords such as "books," journals," "library," "literature," "publications," "references," "archives," and so forth.

Francis Walker's Publications on Treehoppers
2015-08-05
English entomologist Francis Walker (1809–1874) described approximately 361 species of treehoppers; 199 of these are currently considered to represent valid species. Among treehopper enthusiasts, Walker is known for his works on the collections of the British Museum of Natural History (now the Natural History Museum), which holds many, but not all, of his types.

The following links to Walker’s treehopper publications in the Biodiversity Hertitage Library were compiled by Lewis L. Deitz and Matthew S. Wallace in July 2015: Walker 1851a; Walker 1851b; Walker 1852a; Walker 1857a; Walker 1857b; Walker 1858a; Walker 1858b; Walker 1859a; Walker 1860a; Walker 1862a; Walker 1868b; Walker 1868c; [Walker 1870a].

Enter “Francis Walker” in the author search box of the DrMetcalf database to view a full list of Walker’s works in Metcalf’s bibliography of the Auchenorrhyncha.

Note that "Walker 1870a," of Z. P. Metcalf and other authors, is an error for “Walker 1868b” and “Walker 1868c” (here designated to conform to Metcalf’s system of unique identifiers). Based on a published extract, Metcalf catalogued the new genera and species from this work as published in 1870, however, at least two bound copies of volume 10 of the Journal of the Linnean Society of London (Friedrich F. Tippmann's copy at North Carolina State University; also Smithsonian's copy) include an initial page indicating pp. 69-132 were published 7 Aug. 1868 (Walker 1868b), and pp. 133-196 were published 25 Sept. 1868 (Walker 1868c). For purposes of nomenclature, the new taxa included were thus described in 1868 (not 1870).

Carl Stål's Publications on Treehoppers
2015-08-11
Widely regarded as the leading hemipterist of his time, Swedish entomologist Carl Stål (1833–1878) described extraordinary numbers of treehoppers: 124 genera or subgenera, 199 species, and 1 subspecies. Even though a couple of Stål's genera later proved to be leafhoppers, 84 of his treehopper genera remain valid today, as well as 135 treehopper species, 2 subspecies (1 being a nominotypical subspecies), and his well known family group names Smiliinae, Smilini, and Tragopini. Stål published not only on various groups of Hemiptera, but also on beetles, hymenopterans, and orthopterans.

All of Stål's works on treehoppers listed here are currently in the Biodiversity Hertitage Library:Stål 1853b; Stål 1854b; Stål 1855a; Stål 1858a; Stål 1858b; Stål 1858f; Stål 1859b; Stål 1862b; Stål 1862e; Stål 1863c; Stål 1864a; Stål 1866a; Stål 1866c; Stål 1867a; Stål 1869a; Stål 1869c; Stål 1870c.

The HathiTrust Digital Library includes W. D. Funkhouser’s typewritten English translations of Stål 1864a (Funkhouser’s translation); Stål 1866a (Funkhouser’s translation); Stål 1866c (Funkhouser’s translation); Stål 1869c (Funkhouser’s translation).

Enter “Carl Stal” in the author search box of the DrMetcalf database to view a full list of Stål's works in Metcalf’s bibliography of the Auchenorrhyncha.

L. M. H. Fairmaire’s Publications on Treehoppers
2015-08-18
Although best known for his many contributions on Coleoptera (beetles), French entomologist Léon Marc Herminie Fairmaire (1820-1906) worked also on the insect orders Hemiptera and Hymenoptera. His currently valid treehopper taxa include the genera Acutalis, Aconophora, Darnoides, Horiola, Ophiderma, and Smiliorachis and 177 species.

Fairmaire’s works on treehoppers are available online (either the original or a subsequently reprinted version) as follows: Fairmaire 1845a; Fairmaire 1846a; Fairmaire 1846b; Fairmaire 1846c; Fairmaire 1851a; Fairmaire 1855a [see Fairmaire 1885a, a reprint with the same pagination]; Fairmaire 1885a, [Plates 7-9]; Fairmaire and Signoret 1858a, [Plate XI]; Fairmaire and Signoret 1858b [this reprint of Fairmaire and Signoret 1858a (which see) differs in pagination].

Enter "Fairmaire" in the author search box of the DrMetcalf database to view a full list of Fairmaire’s works in Metcalf’s bibliography of the Auchenorrhyncha.

Good Vibrations Key To Insect Communication
2015-09-02
In its special series "Close Listening: Decoding Nature Through Sound," National Public Radio recently aired Good Vibrations Key To Insect Communication (Christopher Joyce and Bill McQuay, 27 August 2015). Highlighting the work of treehopper researcher Rex Cocroft and tree cricket researcher Laurel Symes, the link allows one to hear the original broadcast along with additional communications of treehoppers and tree crickets. For prior NPR broadcasts on treehopper communication, see our Sounds page and archive of past news: “Societies of Sound in the Amazon” (posted 2013-08-29).

Farewell Helonica, Rest in Peace
2015-09-03
Wallace, M. S. 2015b. Taxonomic changes in the treehopper genera Helonica Ball, Telamona Fitch, and Palonica Ball (Hemiptera: Membracidae; Smiliinae: Telamonini). Zootaxa 4007(2): 251-258.

Telamona projecta Butler. © 2010, by Matthew S. Wallace

Wallace’s recent fingings: (1) placed Helonica as a junior synonym of Telamona ; (2) created the new combination Telamona excelsa (Fairmaire), with the new junior synonym T. unicolor Fitch; (3) reinstated Telamona projecta Butler as valid for the species formerly incorrectly identified as "Helonica excelsa" in the central and eastern United States (see figure, above); and (4) reinstated Palonica albidorsata (Fowler), new combination, as a valid species from Veracruz, Mexico.

As a consequence of these changes, Deitz and Wallace’s 2002 checklist of Nearctic species should be further updated as follows: p. 9, delete Helonica excelsa (Fairmaire 1846); p. 13, insert "Telamona excelsa (Fairmaire 1846): USA: Southwestern States: TX; Central & Eastern States: AL, AR, CT, DE, FL, IA, IL, IN, KS, KY, MA, MD, MI, MN, MO, NC, NE, NJ, NY, OH, OK, PA, TN, WI. CANADA: Central & Eastern Provinces: ON, QC. Fairmaire described this species from Mexico (no locality was specified, so this species may be endemic to the Nearctic or may also occur in Neotropical Mexico)"; insert "Telamona projecta Butler 1877: USA: Central & Eastern States: DE, IA, IL, MD, MO, NJ, NY, SC, TN"; delete Telamona unicolor Fitch 1851. For a prior update, see archive of past news: "Recent Papers" (posted 2015-07-03).

iDigBio Portal for Specimen Data on Treehoppers
2015-09-08
Katja Seltmann alerts us that thousands of specimen records for North American treehoppers are now available online at the iDigBio Portal.  Moreover, label data and images for millions of biological specimens are accessible through Integrated Digitized Biocollections (iDigBio), a resource funded by the National Science Foundation.  Browsing the tutorial is recommended to make searches easier and more productive.  Standard search results for treehopper species include a distribution map and a list of specimen records (view sample record for Carynota mera).

Using the Advance Search with " Membracidae" as the search term, Charles Bartlett found more than 113,000 specimen records, including some data not currently displayed in the standard search.   These detailed records can be downloaded in a csv file sent to the email account one specifies.  The specimen data, which in some cases include “host associations” or “host family,” arrive in a zipped folder that includes several files--the most useful of these, the 'occurrence' file, can be opened in Microsoft Excel.  Because the file structure varied among institutions, persistence and patience may aid in reviewing the data. 

Another approach for retrieving data from iDigBio is iDigBio API, an abstraction layer that allows reuse and mashup of aggregated data without a knowledge of the complexity of the back-end data storage.

William Weeks Fowler's Contributions on Treehoppers and 10 Plates by Edwin Wilson
2015-09-13
William Weeks Fowler (1850-1922), an English entomologist and clergyman, is best known to hemipterists for his contributions to the Biologia Centrali-Americana. Among the many new treehopper taxa that Fowler described, 26 genera, 1 subgenus, 143 species, 4 subspecies, and the tribe Acutalini (Smiliinae) remain valid. Fowler’s sections of the Biologia Centrali-Americana include 10 extraordinary plates depicting treehoppers, all drawn and printed by Edwin Wilson, of Cambridge, England ("Introduction" to Insecta: Rhynchota: Hemiptera-Homoptera, Volume 2, Part 1). The highlighted plate numbers under the thumbnails below link to the corresponding plates in the Smithsonian Institution's copy of this work in the Biodiversity Heritage Library.


Plate 1
(Fowler 1894b). Plate 2 (Fowler 1894b). Plate 3 (Fowler 1894c). Plate 4 (Fowler 1894c). Plate 5 (Fowler 1895a).

Plate 6 (Fowler 1895b). Plate 7 (Fowler 1895d). Plate 8 (Fowler 1895e). Plate 9 (Fowler 1896c). Plate 10 (Fowler 1896d)

All of W. W. Fowler's works on treehoppers listed here are in the public domain and accessible through the Biodiversity Heritage Library. The Biologia Centrali-Americana is also online at Electronic Biologia Centrali-Americana (© 2004, Smithsonian Institution Libraries). To generate a full bibliography of Fowler’s works on Auchenorrhyncha, enter “William Weeks Fowler” in the author search box of the DrMetcalf database. Fowler’s Treehopper Publications: Fowler 1894a; Fowler 1894b, Plate 1, Plate 2; Fowler 1894c, Plate 3, Plate 4; Fowler 1895a, Plate 5; Fowler 1895b, Plate 6; Fowler 1895c; Fowler 1895d, Plate 7; Fowler 1895e, Plate 8; Fowler 1895f; Fowler 1896b; Fowler 1896c, Plate 9; Fowler 1896d, Plate 10; Fowler 1896e; Fowler 1897a, Plate 11 [Cercopidae]; Fowler 1904a [see p. 81, note on Lycoderes and Membracidae]; Fowler 1909a Supplement [updates the preceding parts].

Watch, Listen, Learn
2015-09-30
Delight in the sights and sounds of treehoppers in the wild:

Thornbug Treehopper Mother Defends Offspring from Predatory Stinkbug Nymph (© 2014, by Jennifer Hamel)

Treehoppers Communication (© 2013, by H. Carl Gerhardt)

See also our archive of past news: "Treehopper Videos" (posted 2012-12-13).

“Cicadas' Charismatic Cousins” Featured at BugFest 2015
2015-10-01

Science comedian Brian Malow (left) and Auchenorrhyncha specialist Jason Cryan (center) in the Daily Planet Theater, BugFest 2015. Image © 2015, by Matt Zehr.

The North Carolina Museum of Natural Sciences, Raleigh, N. C., is host to BugFest, the largest celebration of insects anywhere. Year after year, this event attracts tens of thousands of insect enthusiasts. BugFest 2015 was held on 18 September 2015, and the focal insect was the "cicada." Treehopper aficionados will be pleased to learn that Jason Cryan provided balance to the program by highlighting membracids and other charming cicada relatives in his superb talk Cicadas' Charismatic Cousins.

Treehoppers: Under Appreciated or Keystone Taxa?
2015-10-02
Wagner, D. L.; Gagliardi, B. L. 2015a. Hairstreaks (and other insects) feeding at galls, honeydew, extrafloral nectaries, sugar bait, cars, and other routine substrates. American Entomologist 61(3): 160-167.

In their discussion of the importance of non-floral sugar resources (NFSRs) to hairstreak butterflies, moths, and "hordes" of other insects, entomologists Wagner and Gagliardi suggest these non-floral resources may exceed even flowers, as the primary source of carbohydrates for thousands of adult holometabolous insects. Thus, treehoppers and other honeydew-producing Hemiptera possibly play a far greater role in the population dynamics of insects than anyone previously recognized.

We conclude that treehoppers are indeed keystone taxa. In addition to being significant components of many ecosystems around the world, treehoppers increasingly serve as model organisms for studies of ecology, evolution, and behavior, and also bring joy to all who appreciate the beauty and diversity of the natural world. [L. L. Deitz and M. S. Wallace]

Immature Stages of New World Treehoppers: First Installment, Amastrini
2015-10-04
McKamey, S. H.; Wallner, A. M.; Porter, M. J. 2015a. Immatures of the New World treehopper tribe Amastrini (Hemiptera, Membracidae, Smiliinae) with a key to genera. ZooKeys 524: 65-87.

This open access publication is the first in a series intended to provide a guide to identifying the immatures of the New World treehopper genera. Although focused on the tribe Amastrini, this installment reviews prior studies on nymphal membracids in general and provides an overview of the morphological features used to describe them. Here, McKamey et al. provide the first descriptions of the immatures of 8 genera (of 11 total in the tribe), along with an illustrated key to the genera and observations on their biology. With 65 figures (many in color), this work is an essential resource for identifying nymphs of the tribe Amastrini.

Based on its distinct nymphal morphology, Vanduzea laeta nolina Ball is elevated to the rank of species: Vanduzea nolina, new status. Nymphal morphology is also used to support the recognition of Bajulata as a valid genus, distinct from Vanduzea.

We learn from the paper’s literature review that the earliest known study on treehopper immatures seems to have been completed between 1783 and 1794, by H. R. Scheller [published in 1869: Scheller 1869a, plates 8-10]. Nevertheless, even today the immatures of most membracid taxa remain to be described. Thus, we eagerly look forward to future papers by McKamey and associates that will describe the nymphs of additional New World genera and tribes.

Guide to Treehopper Workers of the 1700s and Their Publications
2015-10-16
This guide provides access to virtual copies of most of the literature on treehoppers by scholars of the 1700s, including their further works into the 1800s. One exception is Pierre A. Latreille, whose 1796a work is listed here, but not his 24 additional references on treehoppers published during the 1800s. Also excluded here are workers, such as H. R. Scheller, whose first publication appeared after 1799. The earliest work listed here is Linnaeus’s (1746a: p. 202) description of his species number 641, subsequently named Centrotus cornutus Linnaeus, 1758. Owing to modern technology and awe-inspiring international collaboration, this collection of rare and priceless publications is at your fingertips.

Early contributions on treehoppers were sometimes published over a period of two or more years and subsequently bound into a single volume with a title page and introductory materials. Because of the principle of priority among taxonomic names, it is essential to establish the actual date of publication for each section of such works that describe new taxa (the section date may differ from that on the title page). Furthermore, the sequence of sections and placement of plates may vary among various bound copies of a work. In the absence of photography, illustrations were frequently produced from copper engravings (plates), and some were individually hand-colored, each plate being a unique work of art.

Selected early natural history plates with figures of treehoppers: 1, Coquebert 1799a: Pl. 8, Fig. 1; 2 , Coquebert 1799a: Pl. 9, Fig. 5; 3, Coquebert 1801a: Pl. 18, Figs. 1-10; 4, Coquebert 1804a: Pl. 21, Fig. 5; 5, Houttuyn 1766a: Pl. 81, Fig. 2; 6, Linnaeus 1761a: Pl. 2, Fig. 279; 7, Panzer 1798a: Pl. 50.19, Membracis cornutus; 8, Panzer 1798a: Pl. 50.20, Membracis genistae; 9, Donovan 1794a: Pl. 83; 10, Donovan 1820d: Hemiptera Pl. 3, C. spinosa, C. taurus; 11, Stoll 1788a: Pl. [28], Figs. 163, A, B, 165, D, 166, E, 169, H; 12, Sulzer 1761a: Pl. 10, Fig. 63; 13, Sulzer 1776a: Pl. 9, Fig. 6.

1 2 3 4
5 6 7 8 9
10 11 12 13

Z. P. Metcalf took care in establishing publication dates, and we follow his dates unless convincing evidence has come to our attention indicating a needed correction. In this regard, during the compilation of the present guide, we discovered Index Novus Litteraturae Entomologicae, an updated bibliography of the literature on entomology from the beginning until 1863, which appears to be a useful resource for verifying publication dates. To generate full citations for nearly all works published on Auchenorrhyncha during the 1700s, enter "17??" in the date search box of DrMetcalf.

The names of early authors often vary in spelling depending upon the language of the publication, thus variants are given in square brackets in the following list of Treehopper Workers of the 1700s: James [Jacque] Barbut; Johann Beckmann [Iohanne Beckmanno]; Nikolaus Joseph [Nicolaus Josephus] Brahm; Antoine Jean [Antoine-Jean, Antonius-Joannes, Antoine-Jean-Joseph, Jean Antoine] Coquebert [Coquebert de Montbret]; Charles [Caroli] De Geer; Charles [Carolo] Joseph De Villers [de Villers]; Edward Donovan; Johann [Iohannes, Jean, Johan, Johann, Johannes] Christian [Christianus, Chrtien] Fabricius; Johann Reinhold Forster [Joanne Reinholdo Forstero]; Antoine François [Francois, Francoise] Fourcroy [de Fourcroy]; Johann Kaspar [Caspar] Füssli [Fuessli, Fuesslins, Fuessly]; Étienne [Etienne] Louis Geoffroy; Michel Esprit Giorna; Johann Friedrich Gmelin; Johann August Ephraim Goeze; Laurens [Laurentius] Theodorus Gronovius; P. Höslin; Martin [Martinus] Houttuyn; Pierre André [Andre] Latreille; Anton August Heinrich Lichtenstein; Carolus Linnaeus [Carl von Linné]; Christian Friedrich Ludwig; Maria [María, Marie, Anna Maria] Sybilla [Sybille] Merian [de Merian]; Otto Frederik [Friedrich] Müller [Mueller]; Philipp Ludwig Statius Müller [Mueller]; Guillaume Antoine Olivier; Peter Simon Pallas; Georg Wolfgang Franz Panzer; Vincenzo Petagna; Erich [Erik] Pontoppidan; Anders Jåhan [Jahan] Retzius; Johann Jacob Römer [Roemer]; Pietro Rossi; Jacob [Jakob] Christian Schäffer [Schaeffer]; Franz von Paula [de Paula] von Schrank [Schrank]; Johann Christian Daniel von Schreber; Giovanni [Johannes, Ioannis] Antonio [Antonii] Scopoli; Caspar Stoll; Johann [Johan] Heinrich Sulzer; Carl Peter [Caroli Petri, Carolo Petro] Thunberg [Thvnberg]; Friedrich [Frederico] Weber; Thomas Pattinson Yeats; Johann Jacob [Iohannes I.] Zschach [Zschachii] and Nathanael Gottfried Leske [Leskeanum].

Publications on Treehoppers by Workers of the 1700s: Barbut 1783a, pl. 9 ; Beckmann 1772a; Brahm 1790a; Coquebert 1799a, pls. 8-9; Coquebert 1801a, pl. 18; Coquebert 1804a, pl. 21; De Geer 1773a [see pp. 158-328 and pl. 11, Fig. 23, image incomplete]; De Geer 1778a, [see pp. 158-228, Cigales], [plate 45]; De Villers 1789a; Donovan 1794a, pl. 83; Donovan 1820a, [see Cicada]; Donovan 1820d, Hemiptera pl. 3, C. spinosa, C. taurus; Fabricius 1775a; Fabricius 1777a; Fabricius 1781a; Fabricius 1787a; Fabricius 1794a; Fabricius 1798a; Fabricius 1803a; Forster 1771a; Fourcroy 1785a; Füssli 1775a; Geoffroy 1762a, pl. 9 ; Geoffroy 1764a; Geoffroy 1799a; Giorna 1791a; Gmelin 1789a; Goeze 1778a; Goeze 1780a [needed]; Gronovius 1764a, [see ICZN Opinion 261: rejected for nomenclatural purposes]; Höslin 1782a [needed]; Houttuyn 1766a, pl. 81; Latreille 1796a [24 treehopper publications in the 1800s are not listed here]; Lichtenstein 1796a, [see ICZN (1995) Opinion 1820, Bulletin Zoological Nomenclature 51: 283-285]; Linnaeus 1746a [Laurentii Slavii printing], [Lugundi Batavorum printing]; Linnaeus 1758a; Linnaeus 1761a, pl. 2; Linnaeus 1764a; Linnaeus 1767a [subsequent reprints of 1758a not listed here]; Ludwig 1799a; Merian 1771c; Müller, O. 1776a; Müller, P. L. S. 1774a [needed]; Olivier 1789a; Olivier 1792a; Olivier 1797a, pl. 109; Pallas 1766a, pl. 14; Pallas 1772a, pl. 1; Panzer 1798a, pls. 50.19-50.20; Panzer 1802a [needed]; Panzer 1804a; Petagna 1787a; Petagna 1792a, pl. 9; Petagna 1808a [needed]; Petagna 1820a [needed]; Pontoppidan 1762a [see p. 680, Hemiptera, and pl. 29 Cicada cornuta]; Retzius 1783a; Römer 1789a, pl. 9; Rossi 1792a, [see pp. 212-220, Rhyngota]; Rossi 1794a, [see p. 50, Ryngota]; Rossi 1807a, [see pp. 341-352, Rhyngota]; Schaeffer 1767a, pl. 96; Scheller 1869, [plates]; von Schrank 1781a; von Schrank 1801a; von Schreber 1759a; Scopoli 1763a; Scopoli 1777a; Stoll 1781a; Stoll 1788a, pls. 1-2, pls. 5-6, pl. 8, pls. 10-11, pls. 15-[19], pl. 21, pl. [28]; Stoll 1792a [German translation of 1788a]; Sulzer 1761a, [see pp. 94-98, 24-25 and pl. 10]; Sulzer 1776a, pl. 9; Thunberg 1789a; Thunberg 1791a; Thunberg 1822a [needed]; Weber 1795a, [see pp. 144-148, Rhyngota; Weber 1801a, [see pp. 112-115, Rhyngota]; Yeats 1773a; Zschach and Leske 1788a, [see pp. 115-117, Ryngota].

Please contact Lewis Deitz if you know of other works mentioning treehoppers printed prior to 1800 or if you can provide links to online copies of works marked here as “needed.”

Units Used to Measure Insects Prior to the Metric System
2015-10-23
Be grateful that millimeters have largely replaced “lines” and “lunules” as units to measure treehoppers. By the way, a “lunule” is the white crescent-shaped area at the base of one’s fingernail--lunules vary in size and may even be absent in some individuals. More frequently, early entomological workers used “lines” as a unit of measurement, but the definition of a line varied among the English, French (ligne), and Germans (linie), and, in some instances, the definition varied even within a country. The Website www.coleoptera.com includes a brief overview of early entomological measurements. To convert some of the more commonly used "lines" to millimeters, see “Notes on the line as a unit of measurement,” p. 11, from von Hayek, 1973, Bulletin of the British Museum (Natural History), Entomology Supplement 20: 1-309.

Treehopper Publications of E. P. Van Duzee and L. B. Woodruff
2015-11-09
Most works on treehoppers by American entomologists Edward Payson Van Duzee (1861-1940) and Lewis Bartholomew Woodruff (1868-1925) are readily available online through the Biodiversity Heritage Library (BHL). Two of Woodruff’s papers (1923a, 1924a), not currently available at BHL, are accessible through the digital library JSTOR (see Options for Access). For detailed citations of the works by either author, enter his last name in the author search box of DrMetcalf. We note that "Van Duzee 1890a,” which seems to treat only extant leafhoppers, was apparently listed in Metcalf and Wade’s 1963 bibliography on treehoppers and fossil Auchenorrhyncha in error.

Hemipterist Edward P. Van Duzee described 3 treehopper genera (Idioderma, Tylocentrus, and Xantholobus), 43 species (36 now valid, including 2 that are also nominotypical subspecies) and 3 subspecies (2 now valid subspecies, 1 invalid). The California Academy of Sciences, where Van Duzee served as curator (1916-1940), holds his collection of Hemiptera (164,442 specimens). Van Duzee’s 1908a plates 1 and 2, below, show simple line drawings of representative North American treehoppers.



Van Duzee’s Treehopper Publications: Van Duzee 1889b; Van Duzee 1890c; Van Duzee 1892d; Van Duzee 1893b; Van Duzee 1894e; Van Duzee 1895e; Van Duzee 1905a; Van Duzee 1906a; Van Duzee 1907a; Van Duzee 1908a; Van Duzee 1908b; Van Duzee 1909a; Van Duzee 1909c; Van Duzee 1912a; Van Duzee 1912b; Van Duzee 1914a; Van Duzee 1914c; Van Duzee 1916a; Van Duzee 1916b; Van Duzee 1917a; Van Duzee 1917b; Van Duzee 1923a; Van Duzee 1925b.

Lewis B. Woodruff authored 18 treehopper species (16 now valid) and 2 subspecies (1 now elevated to species, 1 invalid), all belonging to the tribe Smiliini. See our news posting of 2012-09-14 [archive of past news], for further information on Woodruff, who at one time had a desk at the American Museum Natural History. The following plates are from Woodruff 1915a (pl. 4, figs. 1-5, now Atymna helena); fig. 6, now A. querci) and 1919a (pl. 23; Ophiderma spp. [Woodruff’s captions include some junior synonyms]).



Woodruff’s Treehopper Publications: Woodruff 1915a; Woodruff 1919a; Woodruff 1920a; Woodruff 1923a; Woodruff 1924a.

[Compiled by Matthew S. Wallace and Lewis L. Deitz, 9 November 2015]

Updates to Treehoppers Team Members and Collaborators
2015-11-19
We welcome graduate student Camilo Flórez-Valencia as the newest member of the Treehoppers "Taxa Content Team." A native of Colombia, Camilo is currently working towards his Master’s degree with Brazilian treehopper researcher Olivia Evangelista. One of Camilo's photos, a Colombian Anobilia, is shown below.

New collaborators include entomologists René Limoges and Stéphane Le Tirant (Montreal Insectarium, Montréal, Québec, Canada). Two of René’s recent treehopper photographs from Mindo, Ecuador, are featured below. See our Acknowledgments and archive of past news (posting of 2015-05-12), for further listings of Team Members and Collaborators.

1, Anobilia nigra, adult with nymphs and ant; 2, Alchisme adult with nymphs; 3, Aconophora female with egg mass. Image copyrights: 1, © 2015, by Camilo Flórez-Valencia; 2 and 3 © 2015, by René Limoges.

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New State Records of Treehoppers for Pennsylvania
2015-12-29
Barringer, L. E. 2015a. Occurrence of treehopper (Hemiptera: Membracidae) bycatch on purple panel traps and Lindgren funnel traps in Pennsylvania, with new state records. Great Lakes Entomologist 48(3-4): 172-185.

Surveys incorporating Lindgren funnel traps and purple panel traps in Pennsylvania (2009-2013) captured 57 native treehopper species, including four not previously reported for the state: Heliria gibberata Ball 1925, Palonica pyramidata (Uhler 1877), Telamona projecta Butler 1877, and Telamona westcotti Goding 1893. Barringer’s work suggests that, with proper site selection, Lindgren funnel traps may be especially useful in monitoring the seasonal distribution of treehoppers, particularly species in the tribes Smiliini and Telamonini.

E. D. Ball’s Publications on Treehoppers
2016-01-19
American entomologist Elmer Darwin Ball (1870–1943) was a specialist on treehoppers and other Auchenorrhyncha. Born in Vermont, he graduated from Iowa State College (B.S., M.S.) and Ohio State University (Ph.D.). Working briefly as a school teacher, Ball had an energetic career with appointments across the U.S.: Albion Seminary (assistant principal), Iowa State College (taught entomology and zoology; later, department head), Colorado Agricultural College (taught entomology and zoology), Utah Agricultural College (professor), the United States Department of Agriculture, the Florida State Plant Board, and the University of Arizona (dean of the College of Agriculture; director of the Agricultural Experiment Station). Moreover, he also served as the State Entomologist of Wisconsin and as the Assistant Secretary of Agriculture. Ball’s extensive collection is now held by the National Museum of Natural History, Washington, D.C.

Regarding treehoppers, Ball described one subfamily (Stictocephalinae, now a junior synonym of the tribe Ceresini), 6 new genera (Bajulata, Bryantopsis, Grandolobus, Helonica, Palonica, and Telonaca–all remain valid), 58 new species (52 remain valid--3 are now also nominotypical subspecies), and 13 subspecies (all remain valid, including 1 elevated to species).

Plate 1 from Ball 1903a shows 9 new species he described in the genus Telamona: Fig. 1, T. extrema; 2, T. obsoleta [= T. westcotti]; 3, T. viridia [now in Palonica]; 4, T. ehrhorni [= Heliria sinuata]; 5, T. compacta; 6, T. decorata; 7, T. pruinosa [= T. collina]; 8, T. brevis [= Telamonanthe pulchella]; 9, T. pulchella [now in Telamonanthe].



Publications of E. D. Ball. All but three of Ball’s works on treehoppers are currently available online through the Biodiversity Heritage Library: Ball 1903a; Ball 1905a; Ball 1915b; Ball 1918a; Ball 1920b; Ball 1925a; Ball 1931f [needed (monograph of the tribe Telamonini)]; Ball 1932b [needed (food plants of various groups)]; Ball 1932c; Ball 1933a; Ball 1937d; Osborn and Ball 1897a [needed (hemipteran fauna of Iowa)].

[Compiled by Matthew S. Wallace and Lewis L. Deitz]

2015 Stamps Feature Ecuadorian Treehoppers
2016-02-16
An Ecuadorian issue of stamps entitled "Membrácidos del Ecuador" features stunning images of treehoppers from Ecuador, taken by nature photographer Milan Kozánek of Bratislava, Slovakia. Eight species are represented including members of the Neotropical genera: Antianthe, Cyphonia, Guayaquila, Heteronotus, Membracis, Stegaspis, Thuris, and Tritropidia. For information on other "Treehoppers on Stamps", see our archive of past news dated 2014-02-25. Many thanks to Milan for sharing his wonderful treehopper images with the world and for bringing these stamps to our attention. We also thank Matt Bertone for assistance in preparing the figure below.


Immatures and Nomenclatural Changes in Thuridini
2016-02-18
McKamey, S. H.; Porter, M. J. 2016a. First immature of the New World treehopper tribe Thuridini (Hemiptera, Membracidae, Smiliinae) with a new synonym, a new combination, and a new country record. ZooKeys 557: 85-91. [available online]

This work adds Ecuador and Venezuela to the distribution of the genus Thuris (previously known from Brazil and Peru) and describes the late instar nymphs of T. binodosus (Goding), new combination [with T. depressus Sakakibara as its junior synonym]. The genus may be subsocial based on the observation of an aggregation consisting of four late instars along with four adults. Both nymphs and adults are illustrated.

© 2012, by Milan Kozánek. Adult of Thuris binodosus (Goding).

Excluded from Treehopperdom
2016-02-24
At least 18 genera described as treehoppers have since been reassigned to various other insect groups ranging from pygmy grasshoppers to froghoppers, lacebugs, leafhoppers, and various groups of extinct Hemiptera. An example is the Afrotropical pygmy grasshopper genus Xerophyllum Fairmaire (1846a: pl. 4, figs. 10-12 [see below]) that superficially resembles members of the Neotropical treehopper genus Membracis. Thus, some genera listed in early catalogues of the family Membracidae may be missing in more recent listings for the group. Additionally, the genus Eteonus, although not initially described as a treehopper, was inadvertently listed as a membracid by Neave 1939b. The present summary directs users to the current status and placement of these bewildering genera that may seem to have vanished from treehopperdom.


Left, Fairmaire 1846a: pl. 4. Right, details of figs. 10-12, Xerophyllum.

Absoluta Bekker-Migdisova 1962a: 92. Type species: Absoluta distincta Bekker-Migdisova. Status: valid genus (extinct Hemiptera: Coleorrhyncha). [McKamey 1998a].
Bohemania Stål 1855a: 97. Type species: Bohemania sobrina Stål. Status: junior synonym of Coloborrhis (Hemiptera: Cicadellidae: Ulopinae: Ulopini). [McKamey 1998a].
Cercopyllis Scudder 1890b: 470-472. Type species: Cercopyllis justiciae Scudder. Status: valid genus (extinct Insecta, incertae sedis). [McKamey 1998a].
Ceresopsis Bekker-Migdisova 1958a: 66. Type species: Ceresopsis costalis Bekker-Migdisova. Status: valid genus (extinct Hemiptera: Heteroptera). [McKamey 1998a].
Coloborrhis Germar 1836a: 73. Type species: Coloborrhis corticina Germar. Status: valid genus (Hemiptera: Cicadellidae: Ulopinae: Ulopini). [Funkhouser 1927f; Evans 1954a; Oman, Knight, and Nielson 1990a].
Darniopsis Bekker-Migdisova 1958a: 65. Type species: Darniopsis tragopea Bekker-Migdisova. Status: valid genus (extinct Hemiptera: Heteroptera). [McKamey 1998a].
Eteoneus Distant 1903c: 129. Type species: Serenfkia dilaia Distant. Status: valid genus (Hemiptera: Heteroptera: Tingidae (listed as a membracid in error by Neave 1939b)]. [Neave 1939b; Drake and Ruhoff 1960]
Eustollia Goding 1926a: 105. Type species: Cicada jubata Goding, a junior synonym of Type species: Proconia marmorata (Fabricius). Status: junior synonym of Proconia (Hemiptera: Cicadellidae: Cicadellinae: Proconiini). [Funkhouser 1927f; Oman, Knight, and Nielson 1990a].
Evryprosopum Stål 1853b: 267. Type species: Bohemania sobrina Stål. Status: junior synonym of Coloborrhis (Hemiptera: Cicadellidae: Ulopinae: Ulopini: Coloborrhis). [Funkhouser 1927f; Oman, Knight, and Nielson 1990a].
Euryprosopum Stål 1858a: 234. Status: error for Evryprosopum, a junior synonym of Coloborrhis (Hemiptera: Cicadellidae: Ulopinae: Ulopini: Coloborrhis). [Funkhouser 1927f; Oman, Knight, and Nielson 1990a].
Maguviopsis Bekker-Migdisova 1953a: 463. Type species: Maguviopsis kotchnevi Bekker-Migdisova. Status: valid genus (extinct Hemiptera: Cicadomorpha, incertae sedis). [McKamey 1998a].
Membracites Heer 1853b: 92. Type species: Membracites cristatus Heer. Status: valid genus (extinct Pterygota, incertae sedis). [McKamey 1998a].
Minuta Bekker-Migdisova 1958a: 64. Type species: Minuta heteropterata Bekker-Migdisova. Status: valid genus (extinct Hemiptera: Coleorrhyncha). [McKamey 1998a].
Narnia Walker 1870a: 192. Type species: Narnia rastrata Walker 1870a. Status: junior synonym of Machaerota (Hemiptera: Machaerotidae: Machaerotinae: Machaerotini); also a junior homonym of Narnia Stål, 1862 (Heteroptera: Coreidae). [Walker 1870a; Lallemand 1912a; Funkhouser 1927f; Metcalf 1960b; Wallace and Deitz 2004a (incorrectly listed as unplaced within Centrotinae)].
Porcorhinus Goding 1903a: 38. Type species: Porcorhinus mastersi Goding. Status: valid genus (Hemiptera: Cicadellidae: Ledrinae: Ledrini). [Funkhouser 1927f; Oman, Knight, and Nielson 1990a; Jones and Deitz 2009a].
Prolystra Oppenheim 1888a: 228. Type species: Prolystra lithographica Oppenheim. Status: valid genus (extinct Hemiptera: 'Homoptera', incertae sedis). [McKamey 1998a].
Sphongophoriella Bekker-Migdisova 1958a: 63. Type species: Sphongophoriella reticulata Bekker-Migdisova. Status: valid genus (extinct Hemiptera: Heteroptera). [McKamey 1998a].
Tegulicicada Lin 1986a: 62. Type species: Tegulicicada plana Lin. Status: valid genus (extinct Hemiptera: Coleorrhyncha: Karabasiidae). [McKamey 1998a].
Williamsiana Goding 1926a: 103. Type species: Williamsiana ferruginosa Goding, a junior synonym of Zyzzogeton haenschi Breddin. Status: junior synonym of Zyzzogeton (Hemiptera: Cicadellidae: Cicadellinae: Proconiini). [Funkhouser 1927f; Oman, Knight, and Nielson 1990a]
Xerophyllum Fairmaire 1846a: 242. Type species: Xerophyllum servillei Fairmaire. Status: valid genus (Orthoptera: Tetrigidae). [Funkhouser 1927f; Devriese 1999 (Belgian J. Entomol. 1:35)]

Compiled by Lewis L. Deitz, with thanks to Dmitry Dmitriev for pointing out the homonymy of Narnia.

More Ecuadorian Treehopper Stamps
2016-03-03
Paul Freytag alerted us to another issue of stamps in the series "Membrácidos del Ecuador." This issue includes images of members of the genera: Adippe, Alchisme, Cladonota, Cyphonia, and Membracis, all taken by nature photographer Milan Kozánek.



For other treehoppers on stamps, see our archive of past news dated 2014-02-25 and 2016-02-16. We thank Matt Bertone for assistance in preparing the figure above.

Update on Treehopper Types at North Carolina State University
2016-03-07
Since our posting of 2014-02-03 titled "Membracid Types at North Carolina State University," links to NCSU's Insect Museum have been changed and the collection now holds type material of 49 membracid species. For primary types, the listing includes the original binomen, author(s), and literature citation, along with information quoted from the data labels. For some species, images of the type and its labels are available online, as shown below for the holotype of Aconophora elongata (photos Copyright © 2013, by Robert L. Blinn).



Aconophora elongata Dietrich In Dietrich & Deitz 1991. N. C. Agric. Res. Ser. Tech. Bull. 293:112.
Holotype: "PERU/Chancha/mayo; 21-vii-60/J. Salazar; on/Leguminosae; DIETRICH/RESEARCH/4-84-373c; HOLOTYPE/Aconophora/elongata/Dietrich"
Paratypes, 8: Peru

Gonoconophora Awareness Month
2016-04-01
This month we focus on the little known Brazilian genus Gonoconophora Fonseca 1950, which is of uncertain position within the subfamily Darninae (Deitz 1975). The genus is known only from the holotype of its type species – G. funkhouseri Fonseca – which is apparently lost (Evangelista, Santos, and Lamas, 2014a, Zootaxa 3895(1): 1–30). Consequently, everything we known of Gonoconophora is based on its original description by José Pinto Fonseca, 1950a, "Contribuição para o conhecimento dos membracídeos neotrópicos (V)" [Contribution to the knowledge of the Neotropical Membracidae (V)] (Arquivos do Instituto Biológico, São Paulo, 19: 111-116, Plate 8). Except as noted otherwise, the contents of the Arquivos do Instituto Biológico, São Paulo, are licensed under Creative Commons Attribution-NonCommercial 4.0 International (CC BY-NC 4.0). We here reproduce Fonseca’s 1950 original illustrations along with a generic diagnosis translated and modified from Fonseca’s original. Our hope is that increased awareness of Gonoconophora might lead to the discovery of additional material, either in collections or by stimulating efforts to find this treehopper in the wild.



Gonoconophora funkhouseri Fonseca, female holotype. A, lateral aspect; B, dorsal aspect, C, anterior aspect. ©1950, by Arquivos do Instituto Biológico, São Paulo.

Diagnosis: Head transverse, more than four times broader than high. Forewing with five apical cells, in repose, half its width concealed by pronotum. Hind wing hyaline with four apical cells. Pronotum: humeral angles prominent, nearly acute apically; frontal horn directed anterodorsally, laterally compressed, without lateral keels; posterior process well developed, strongly spine-shaped apically, slightly surpassing apex of forewings in length. Body glabrous. Measurements (female holotype): overall length of pronotum 13mm (apex of anterior process to apex of anterior process); length from base of head to wing tip 8mm; between humeral angles 2mm.

Holotype (female): Onda Verde, State of São Paulo, Brazil, 20 January 1946, Frederico Lane (originally deposited in the Collection of the Department of Zoology, State of São Paulo, Brazil).

Prepared by Lewis L. Deitz and Olivia Evangelista.

24th Annual Treehopper Aggregation: 3-5 June 2016, Little Orleans, Maryland
2016-04-04
Join the 2016 Treehopper Gathering, set for 3 to 5 June (arrive Friday, depart Sunday), at the Little Orleans Campground, Little Orleans, Maryland. Nicknamed “treehopper camp” by some, this casual event provides a pleasant opportunity to meet and collect with others passionate about treehoppers and related insects in scenic western Maryland. Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening.

To reserve a campsite with our group or to obtain further information, contact Charles Bartlett. Advanced registration with Charles is essential to assure that you have a campsite.

Bring your collecting vials, insect nets, unidentified specimens, cameras, and bathing suits!

New Tex-Mex Treehopper Celebrates M. S. Wallace and Selena
2016-04-26
Music lovers and entomologists are abuzz with news of Selenacentrus wallacei, new treehopper genus and species, recently described from Texas and northern Mexico by Brendan O. Morris and Christopher H. Dietrich (2016, "Hidden in plain sight: a remarkable new genus of Nearctic treehoppers (Hemiptera: Membracidae)," Annals of the Entomological Society of America [advance access published 29 March 2016; DOI: http://dx.doi.org/10.1093/aesa/saw008]. The new genus name honors Selena Quintanilla, beloved Queen of Tenjano [or Tex-Mex] music. The species name honors esteemed treehopper specialist Matthew S. Wallace. Additional work is needed to clarify the phylogenetic position of this new membracid relative to other taxa within the subfamily Centrotinae. The treehopper feeds on a few members of the plant family Fabaceae (Prosopis, Pithecellobium, and Acacia).

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© 2016, by Brendan O. Morris. Selenacentrus wallacei Morris and Dietrich. 1-2, female; 3-4 male.

Read more at EntomologyToday, Science Daily, and Phy.org.

Celebrating 1 May 1885: Z. P. Metcalf’s Birth
2016-05-03
Below we feature a clever plaque presented to Dr. Zeno Payne Metcalf on his 65th birthday by the “Membracids” (friends, faculty, students, and family at NC State University). Among the distinguished signatories is Virginia Wade, who assisted in compiling Metcalf’s invaluable catalogue of the Auchenorrhyncha. Although Metcalf died on 5 January 1956, his legend lives on as documented on the NCSU Libraries’ Website "DrMetcalf: a Resource on Cicadas, Leafhoppers, Planthoppers, Spittlebugs, and Treehoppers." We thank Micou M. Browne (Metcalf’s grandson) for sharing this long forgotten historical plaque with all who treasure treehoppers and Metcalf’s remarkable contributions.

© Copyright 1950, by the Membracids. Facsimile by Matthew A. Bertone.

Funkhouser’s 1951 “Homoptera Fam. Membracidae.” Genera Insectorum, Fascicle 208
2016-05-17
With great pleasure, we provide a Biodiversity Heritage Library link to “Funkhouser 1951a,” William D. Funkhouser’s final and most comprehensive contribution (of many) on treehoppers. Although the classification and nomenclature are out-of-date for many taxa, this remains a valuable reference. Funkhouser’s Plates 1-27 precede fascicle 209 on Diptera by E. Séguy in this online copy.

Short Film Featuring Animated Treehoppers
2016-06-10
Treehopper fans, enjoy this wonderful animation entitled Invisible Nature: Code of the Treehopper, produced for the Cal Academy! It stars Enchenopa and Umbonia, as personified by Rafa Rodriguez and Carol Miles, with some background and recordings provided by Rex Cocroft.

Notes and Photos from the 24th Annual Treehopper Gathering
2016-06-20
Treehopper enthusiasts representing 10 of the 50 States, as well as China, Colombia, and Russia, attended the 24th Annual Treehopper Gathering, 5 to 7 June 2016, Little Orleans, Maryland. Attendees (photo 1, left to right) included: Dmitry Dmitriev, Ashley Kennedy, Charles Bartlett, Andrew Short, Nate Nazdrowicz, Anthony Gonzon, Yang Wang, Anthony Deczynski, Laura Catalina González-Mozo, Matt Wallace, Lawrence Barringer, Dawn Flynn, Vlad Dmitriev, Lewis Deitz, Jason Owens, Mark Rothschild, Kelley Tilmon, Karen McGowan, Kerry McGowan, Stuart McKamey, Hava Amsbury, Dennis Kopp, Eli Wyman, and, in photo 2, Felipe Gonzalez-Mozo (he napped during the group photos).

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2 Photos 1-2 © Copyright 2016, by Dmitry Dmitriev.

Aside from one torrential downpour, the weather was fine. As usual, base camp activities centered around the picnic shelter and ranged from treehopper mounting, sorting, and identification (photo 3 [left to right Laura G.-M., Stu McK., and Yang W.]) to lending and exchanging specimens, trading literature, socializing, playing Frisbee, showing off prized treehopper t-shirts (photo 4, shirt presented to Dennis K. from friends at North Dakota State University, 1990), and enjoying the Saturday evening feast of meats and veggies (photos 5-7: 5, Charles B. grilling steak; 6, Ashley K. grilling veggies; 7, Anthony G. smiling over his fried turkey). An exciting development was that Dmitry Dmitriev (Illinois Natural History Survey, Champaign) will be collaborating with our existing Treehoppers Teams to assist in developing the Treehoppers Database in TaxonWorks. Thus, Dmitry is now officially part of our Taxa Content Team, Nomenclatural Database Team, and References Database Team.

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Photos © Copyright 2016: 3, 5-7 by Dmitry Dmitriev, 4, by Lewis Deitz.

Many participants also had fun collecting (photos 8-11: 8, Dmitry D.; 9, Matt W. with beating sheet and Vlad D. with sweep net; 10, Mark R., Kerry McG., and Karen McG.) and exploring areas around the campground (photo 11, Vlad D. and Yang W. at Fifteen Mile Creek). Notably, Matt W. collected his first nymph of Heliria excelsa, which he reared to the adult stage, thus filling a significant gap in his ongoing research on the nymphs of the tribe Telamonini. Among numerous other specimens collected were Telamona ampelosidis [male] (photo 12), Entylia carinata (13), and a fine-looking nymph of the genus Glossonotus (photo 14).

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© Copyrights 2016, by Dennis Kopp (photo 8), Dmitry Dmitriev (photos 9-11). Photos 12-14, by Stuart McKamey, are NOT IN COPYRIGHT.

Because 2017 will mark our 25th Annual Treehopper Gathering, we will likely extend next year’s gathering by an extra day or two! We hope to announce the final dates 2 or 3 months before the event, most likely it will include the first weekend of June. Already folks are thinking about special t-shirt designs to celebrate our Silver Anniversary.

Publications on New World Treehoppers from 2016
2016-08-09
Benassi, V. L. R. M.; Valente, F. I.; Souza, C. A. S.; Benassi, A. C.; Sakakibara, A. M. 2016a. Biodiversidade e sazonalidade de cigarrinhas (Hemiptera: Membracidae) em cacaueiros. EntomoBrasilis 9(1): 41-46. [available online; in Portuguese, with English abstract].

This study lists and illustrates 30 membracid species associated with Theobroma cacao in Linhares, Espirito Santo State, Brazil. Seasonality data are given for 5 of 8 species confirmed to feed on cacao. Horiola picta was by far the most abundant species.

Creão-Duarte, A. J.; Sakakibara, A. M. 2016a. Two new species of Sundarion (Hemiptera: Membracidae: Darninae) and taxonomic notes on two congeneric taxa. Zoologia 22(1):1-7.

Published online March 2016, this work includes: Sundarion flavopiceum and S. marmoratum n. spp.; S. compressicornis (Fairmaire, 1846), n. comb.; and notes on S. notable de Souza [Evangelista de Souza] and Rothéa.

Flynn, D. J.; Wheeler, A. G., Jr. 2016a. Micrutalis pallens Fowler (Hemiptera: Membracidae): first U.S. records, host-plant association, description of male, and redescription of female. Proceedings of the Entomological Society of Washington 118(3):345-353.

This paper extends the distribution of M. pallens to the Nearctic region, with records on Thurber’s desert honeysuckle (Acanthaceae: Anisacanthus thurberi) in Arizona and New Mexico. Prior reports of this treehopper were all from the Neotropics (Argentina, Guatemala, Guyana, Mexico, Panama, and Venezuela). M. pallens is compared to five other species Micrutalis reported from the Southwest.

Morris, B. O.; Dietrich, C. H. 2016a. Hidden in plain sight: a remarkable new genus of Nearctic treehoppers (Hemiptera: Membracidae). Annals of the Entomological Society of America 109(3): 488-494. [in English, with Spanish abstract].

First published online, this paper describing Selenacentrus wallacei, new genus and species, is now available in the print version of the journal. This treehopper occurs on a few members of the plant family Fabaceae (Prosopis, Pithecellobium, and Acacia) in Texas and northern Mexico.

Pinto, C. F.; Salinas, S.; Flores-Prado, L.; Echeverría, J.; Niemeyer, H. M. 2016a. Sequestration of tropane alkaloids from Brugmansia suaveolens (Solanaceae) by the treehopper Alchisme grossa (Hemiptera: Membracidae). Biochemical Systematics and Ecology 66: 161-165.

Both adult males and females of A. grossa sequester tropane alkaloids from the host B. suaveolens.

Wallace, M. S. 2016a. Telamona barbata Van Duzee 1908, reinstated junior synonym of T. decorata Ball 1903, and their respective lectotype designations (Hemiptera: Membracidae). Proceedings of the Entomological Society of Washington 118(3): 477-479.

Wallace designates, compares, and illustrates the male lectotypes.

Wheeler, A. G., Jr. 2016a. Thelia bimaculata (F.) and Vanduzea arquata (Say) (Hemiptera: Membracidae) in the southern Appalachians: new hosts in Robinia (Fabaceae) and nymphal use of abandoned cavities of the locust twig borer. Proceedings of the Entomological Society of Washington 118(3): 436-449.

Wheeler provides extensive field observations of these membracids in the Appalachians of North Carolina, South Carolina, and Tennessee.

An Alarm Signal to Guard Treehopper Eggs Against Mymarids?
2016-10-28
Miranda, X. 2016a. Egg-guarding behavior of the treehopper Ennya chrysura (Hemiptera: Membracidae): female aggregations, egg parasitism, and a possible substrate-borne alarm signal. Revista de Biologia Tropical 64(3): 1209-1222. [available online]

Substrate-borne signals produced by females of E. chrysura while guarding their egg masses against parasitism may alert neighboring females of an attack, thus lowering the overall rate of parasitism by certain mymarid wasps. Miranda observed that under natural conditions females tend to deposit their eggs close to other egg-guarding females. Also, in the laboratory, females exposed to playbacks of another female’s disturbance signal react more quickly and defensively to a tactile stimulus.

First Treehopper in Niue
2016-11-11
Wallace, M. S.; Anderson, D.; Deitz, L. L. 2016a. Antianthe expansa (Germar): first treehopper (Hemiptera: Membracidae) recorded from Pacific Island Nation of Niue. Proceedings of the Entomological Society of Washington 118(4):650-653.

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Vouchers of Antianthe expansa collected in Niue, 21 January 1996, on tomatoes: 1-2, adult (1, anterior; 2, lateral), 3-4 nymph (3, anterior; 4, lateral).

This work documents Antianthe expansa (Figs. 1-4) on tomatoes in Niue, however, a prior record from Hawaii and an unconfirmed report from Tonga are also discussed. Thus, this New World treehopper is reported not only from the Neotropical and Nearctic regions, but also in the Oceanic region.

Chinese Version of Maruyama's "Treehoppers: Incredible Insects""
2016-12-08
Chung-Ping Lin informs us a Chinese version of Maruyama’s book "Treehoppers: Incredible Insects" is being published this month (December 2016; ISBN:9789864431885). A few Taiwanese species have been added to the earlier edition (see archive of treehopper news: “Maruyama's 2011 book "Treehoppers: Incredible Insects," dated: 2012-08-03).

Photos by Shoyo Sato, New Collaborator
2017-03-07
We here feature six photos by Shoyo Sato (Boston University, Boston, Massachusetts), a new collaborator on our Treehoppers Team. Shoyo captured these and other treehopper images in Ecuador, while working as a teaching fellow with Kelly Swing in late 2016.

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1, Lophyraspis with ant; 2, Notocera; 3, Guayaquila mackameyi with nymphs; 4, Lycoderides phasianus; 5, Phyllotropis cingulata with nymphs and ants; 6, Stegaspis fronditia.

Evidence from Treehoppers (Hemiptera: Membracidae) of Climate Change?
2017-03-28
Disclaimer. The "Officially Approved” abstract, below, is reproduced precisely as received from an "undisclosed source." We are unable to confirm or deny the data and results reported therein. Thus, any opinions, findings, and conclusions or recommendations expressed in this abstract are strictly those of its unknown author(s) and do not necessarily reflect the views of this Website or any person (living or deceased) or any known institution or agency. Readers are free to draw their own conclusions based on the evidence, however, the proposed date of release (1 April 2017) is somewhat suspicious.


Celebrate the 25th Annual Treehopper Aggregation: 1-4 June 2017, Little Orleans, Maryland
2017-04-06
Join us for the 2017 Treehopper Gathering set for 1 June to 4 June (arrive Thursday, depart Sunday), at the Little Orleans Campground, Little Orleans, Maryland. We note that by popular demand, an extra day is added to this year’s aggregation. Located in scenic western Maryland, this event offers a relaxed, casual setting to meet and collect with others passionate about treehoppers and other insects.

Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration and the group meal provided on Saturday evening.

To reserve a campsite with our group or to obtain further information, contact Charles Bartlett. Advanced registration with Charles is essential to assure that you have a campsite.

Bring your collecting and camping gear, camera, unidentified treehopper specimens, and a bathing suit!

World Hemiptera Day, 21 April
2017-04-20
Thierry Bourgoin, Chair of the International Auchenorrhyncha Society, reminds us that World Hemiptera Day, 21 April, is a great time to promote the 15th International Auchenorrhyncha Congress (IAC) and the 10th International Workshop on Leafhoppers and Planthoppers of Economic Importance, 9-15 July 2017, in Mendes, Rio de Janeiro, Brazil--the first IAC in South America.

On 21 April, join our colleagues worldwide in celebrating the rich diversity in hemipteran forms, biologies, behaviors, ecologies, and interactions with human society. We are especially pleased that the World Hemiptera Day logo, created by Jacek Szwedo, features a handsome treehopper.

© 2016, Jacek Szwedo, CC license: by_nc_sa_3_0.

New Research on the Majestic Treehopper Subfamily Heteronotinae
2017-04-25
Evangelitsa, O.; Sakakibara, A. M.; Cryan, J. R.; Urban, J. M. 2017a. A phylogeny of the treehopper subfamily Heteronotinae reveals convergent pronotal traits (Hemiptera: Auchenorrhyncha: Membracidae). Systematic Entomology 42: 410-428.

This work includes a phylogenetic reconstruction of relationships within the Heteronotinae based on DNA nucleotide sequence data from 5 nuclear and 2 mitochondrial genes for an extensive sampling of 9 of 10 genera currently placed in this subfamily. Furthermore, mapping selected morphological traits of the pronotum on the resulting phylogeny suggested repeated acquisitions of similar traits. Many of these pronotal traits apparently arose independently within the genus Heteronotus. Overall, the data support the monophyly of the subfamily (excluding Darnoides) and five of its included genera, but also suggest the existence of new undescribed genera as well a species complex with Heteronotus delineatus. At the species level, one new combination is proposed and one original combination is reinstated.

Festschrift Celebrating the Career of Albino Morimasa Sakakibara: Zootaxa 4281(1)
2017-07-10
Just in time for the 15th International Auchenorrhyncha Congress in Rio de Janeiro, Brazil (9-15 July 2017), a special issue of Zootaxa is hot off the presses to celebrate the prolific and impressive career of esteemed Brazilian hemipterist Albino M. Sakakibara. This Festschrift (dated 22 June 2017) includes 23 original taxonomic works in honor of Dr. Sakakibara. The Foreword from the Editors (Evangelista, Takiya, and Dietrich 2017b) presents an overview of Sakakibara's extraordinary contributions on the Auchenorrhyncha. It includes a bibliography of his works and a list of 33 patronyms dedicated to him (see, for example, representatives of the treehopper genus Sakakibarella, below). We agree with the editors who conclude that A. M. Sakakibara "will inspire generations of young taxonomists to come."

Evangelista, O.; Takiya, D. M.; Dietrich, C. H. 2017b. The art and science of describing nature's surrealists: Foreword from the Editors. Zootaxa 4281(1): 5-21. [in Evangelista, O.; Takiya, D. M.; Dietrich, C. H. (eds.). 2017a. The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara.] This editorial foreword is available online.

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Figures 1-2. 1, Sakakibarella elongate Creão-Duarte; 2, S. seminigra Creão-Duarte. (Images © 2006, by Albino M. Sakakibara).

Dr. Sakakibara is especially well known for his many outstanding works on Neotropical treehoppers as documented in the bibliography of his works up to April 2017 (Evangelista et al. 2017a, pp. 17-21). Eight original taxonomic publications in the 2017 Festschrift treat treehoppers:

Flórez-V., C.; Evangelista, O. 2017a. New species in the treehopper genus Bocydium Latreille, with description of nymphal stages and observations on their natural history. Zootaxa 4281: 22-57. [in The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara] This work presents an illustrated key to the 20 species of Bocyidium, including 6 n. spp. described by the first author: B. bilobum, B. hadronotum, B. mae, B. sakakibarai, B. sanmiguelense and B. tatamaense; also B. cubitale Richter, 1954, is placed a junior synonym of B. bulliferum Goding, 1930.

Creão-Duarte, A. J.; Cabral, V. A.; Lourenço, A. 2017a. A new species of Lycoderes Sakakibara [sic] (Hemiptera, Membracidae, Stegaspidinae) from Brazil. Zootaxa 4281(1): 58-62. [in The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara] Lycoderes albinoi, n. sp., is described and illustrated--thus 18 valid species are recognized for the genus Lycoderes Germar [sensu Sakakibara 2013a].

Dietrich, C, H. 2017a. A new treehopper genus and species from Puerto Rico (Hemiptera: Membracidae: Stegaspidinae) with notes on other Caribbean Membracidae. Zootaxa 4281(1): 63-69. [in The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara] Bortinctolania, n. gen., and B. sakakibarai, n. sp., are described and placed as incertae sedis within the subfamily Stegaspidinae. A key is provided to those Caribbean genera that lack a posterior pronotal process and are placed incertae sedis either within Membracidae (genus Togotolania) or within Stegaspidinae (3 genera).

Gaiani, M. A. 2017a. Problematode: an enigmatic new genus of Membracidae (Hemiptera with two new species from Venezuela. [in The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara] Problematode, n. gen. (incertae sedis within the family Membracidae), is described to include P. sakakibarai and P. inerme, n. spp., from the Coastal Montane Forests of Venezuela.

Creão-Duarte, A. J.; Rothéa, R. R. A. D.; Lourenço, A.; Cabral, V. A.; Evangelista, O. 2017a. Two rema[r]kable new species of Notocera Amyot and Serville, 1843 (Hemiptera, Membracidae, Hypsoprorini) from the Brazilian Caatinga. Zootaxa 4281(1): 77-89. [in The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara] This work describes and illustrates Notocera colavitei Creão-Duarte and Rothéa, n. sp., and N. sakakibarai Creão-Duarte and Lourenço, n. sp.

Camacho, L. F.; Flórez-V, C.; Evangelista, O. 2017a. Notes on the genus Sakakibarella Creão-Duarte with description of three new species (Membracidae: Membracinae: Hoplophorionini). Zootaxa 4281(1): 90-107. [in The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara] This work presents a key to the 7 species of Sakakibarella, including 3 n. spp. described by Flórez-V and Camacho from Colombia: S. albinoi, S. crassa, S. santaceciliae.

Gonzalez-Mozo, L.; McKamey, S. H.; Ware, J. L.; Hamilton, G. 2017a Two new species of Darnini (Hemiptera: Membracidae) from Colombia and Peru. Zootaxa 4281(1): 108-114. [in The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara] Cyphotes colombiensis, n. sp., from Colombia, and Hypheodana sakakibarai, n. sp., from Peru, are described and illustrated.

McKamey, S. H. 2017a. Two new species of unusual Ceresini (Hemiptera: Membracidae: Smiliinae). Zootaxa 4281(1): 115-119. [in The art and science of describing nature's surrealists: Festschrift Celebrating the Career of Albino Morimasa Sakakibara] Cyphonia sakakibarai, n. sp., from Brazil, and Poppea unispina, n. sp., from Costa Rica, are described and illustrated.

25 Years of Membrasketeers: The Silver Anniversary Aggregation (1-4 June 2017)
2017-07-24
The early June weather at Little Orleans, Maryland, was perfect for the Silver Anniversary of the Annual Treehopper Gathering. Participants (Fig.1) celebrated by feasting, socializing, collecting insects for research or fun (Fig. 2), swimming, hiking, and other activities. A celebratory T-shirt (Figs. 3-4), designed by Matt Bertone and Kelley Tilmon to commemorate the occasion, lists scores of dedicated attendees from prior years, including two (Bartlett and Deitz) who attended all 25 gatherings.

Fig. 1. The 40 attendees for 2017 included (left to right: b= back; f= front): Hova Amsbury, Eli Wyman, bAnthony T. Gonzon, Tyler Hagerty, bAndrew Short, Abby Clarke, bJim Liebherr, Ashley Kennedy, fCera Fisher, Devan George, bJamie Zahniser, Becca Robertson, Matt Wallace, bThomas Fisher, Stuart McKamey, Ann Hajek, bSteve Toth, Dawn Flynn (wearing our 2012 T shirt), Jason Owens, Mark Rothschild, bLewis Deitz, Kelley Tilmon, fNate Nazdrowicz (wore the same shirt also in group photos for 2013, 2014, 2015, and 2016), Charles Bartlett, bLawrence Barringer, bAnthony Deczynski, Shawn T. Dash, bBrian Wiegmann, Laura Catalina Gonzalez Mozo (barely visible), Penelope Park held by Yuri Park, fFelipe Gonzalez Mozo, bMicah Fletcher, fDuncan Park, Sarah Kocher, fAlexis Park, bEthan Whitecotton, Ellie (the dog attendee) held by Katrina Dash, and Tiffany Le Ngoc. [Photo © 2017, by Ethan Whitecotton (more E.W. images).]

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Figs. 2-4. 2, Black lights attract the strangest critters (photo © 2017, by Eli S. Wyman). 3-4, Celebratory T-shirt (photos © 2017, by Matthew A. Bertone): 3, front. 4, back.

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Notable finds among treehoppers were needed immatures for projects on the tribe Telamonini (Matt Wallace) and an elusive species in the Enchenopa binotata complex (Mark Rothschild). Also many specimens were identified, swapped, lent, or donated among the attendees. Adding the icing to the cake, Stuart McKamey made available thousands of astonishing high resolution images of primary types of treehopper species held by U.S. National Museum of Natural History, which we plan to add to the Treehoppers Website overtime (Fig. 5-12). For nearly all of the types, images of the anterior, dorsal and lateral aspects are represented, along with photos of the specimen labels. We are grateful to Stuart for sharing these images. As shown below, each image is worth a thousand words.

Figs. 5-12. Lateral aspects of selected holotypes in the tribe Stegaspidini held by the U.S. National Museum of Natural History, Washington, D.C. [Photos by Gary Oulette, NOT_IN_COPYRIGHT (USNM 2017)]. 5, Flexocentrus brunneus Funkhouser 1930 [= F. felinus (Haviland 1925)]. 6, Lycoderes luteus Funkhouser 1940 [now Lycoderides luteus (Funkhouser 1940)]. 7, Lycoderes (Lycoderides) nathanieli Cryan in Cryan and Deitz 1999 [now Lycoderides nathanieli (Cryan in Cryan and Deitz 1999)]. 8, Lycoderes triangulata Funkhouser 1919 [now Lycoderides triangulata (Funkhouser 1919)]. 9, Stegaspis viridis Funkhouser 1915 [= S. bracteata (Fabricius 1787)]. 10, Stylocentrus rubrinigris Funkhouser 1940. 12, Umbelligerus peruviensis Deitz 1975. 12, Umbelligerus stockwelli Flynn 2014.

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Links to Works by Laurens Theodorus Gronovius and Carl Stål
2017-07-26
Under our news item entitled "Guide to Treehopper Workers of the 1700s and Their Publications" (dated 2015-10-16), the work listed as "Gronovius 1762a [needed]" is available online through the Biodiversity Heritage Library, but should read "Gronovius 1764a." This work includes descriptions of various species of "Cicada", including Membracis foliata (as Cicada foliacea), p. 181, but was rejected for nomenclatural purposes see ICZN Opinion 261. An update has also been made in our original news item.

Under our news item "Carl Stål's Publications on Treehoppers" (dated 2015-08-11) a number of broken links are corrected here, as well as in the original news item, which gives further useful information. Stål's works on treehoppers available in the Biodiversity Hertitage Library include: Stål 1853b; Stål 1854b; Stål 1855a; Stål 1858a; Stål 1858b; Stål 1858f; Stål 1859b; Stål 1862b; Stål 1862e; Stål 1863c; Stål 1864a; Stål 1866a; Stål 1866c; Stål 1867a; Stål 1869a; Stål 1869c; Stål 1870c.

Florida Treehoppers: Collaborator Update
2017-07-31
To our listings of collaborators in the Treehoppers Acknowledgments and Archive of Past News, we here add Lyle J. Buss (Entomology and Nematology Department, University of Florida, Gainesville). Lyle provided many excellent photos of Florida treehoppers, including the selection of live shots shown below. Treehoppers team member Mark J. Rothschild identified the species.

On viewing Lyle’s images of Telamona salvini (Figs. 1-3), one entomologist declared: "They have better camouflage than the U.S. Army!"

Figs. 1-9. 1-3, Telamona salvini, a female photographed against different backgrounds. 4, Smilia fasciata female. 5, Cyrtolobus dixianus female. 6, C. tuberosus female. 7, Archasia belfragei, male. 8, Ophiderma definita female. 9, Telonaca alta male.
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Photos © 2017, by University of Florida, Entomology & Nematology Dept. (Lyle J. Buss).

Flynnia, a new genus of Thuridini, and erroneous records of South American members of the tribe Smiliini
2017-08-30
McKamey, S. H. 2017b. The identity of three South American "smiliine " treehoppers (Hemiptera, Membracidae) and related taxonomic changes in Thuridini. ZooKeys 678:65-72. [available online.]

The new genus Flynnia (Membracidae: Smiliinae: Thuridini) is named in honor of treehopper worker Dawn Flynn; its type species--Ophiderma fascipennis Funkhouser (Figs. 1-3)--is known from Bolivia and Ecuador. Thelia planeflava Fairmaire, from Brazil, is placed incertae sedis within the tribe Polyglyptini (Smillinae). Atymna pilosa Funkhouser, from Peru, is placed in the genus Amastris (Smiliinae: Amastrini). Cyrtolobus (Atymna) atromarginata Goding, from Ecuador, is placed in the genus Antianthe (unplaced within Smiliinae).

Figs 1-3. Flynnia fascipennis (Funkhouser), male holotype (USNM). 1, anterior aspect; 2, lateral; 3, dorsal.
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J. A. Ramos Collection of Treehoppers and Other Auchenorrhyncha
2017-08-31
On 21 August 2017, the North Carolina State University Insect Museum received the personal Auchenorrhyncha collection of the late entomologist José A. Ramos (portrait below). Donated to NCSU by his son Dr. Stuart Ramos, this collection includes approximately 50,000 specimens of cicadas, leafhoppers, planthoppers, spittle bugs, and treehoppers from the Caribbean Islands and Central and South America. Dr. J. A. Ramos, an alumnus of NC State, earned both his M.S. (1945) and Ph.D. (1956) at State, working under the direction of Zeno P. Metcalf and later Clyde F. Smith. The J. A. Ramos collection of Auchenorrhyncha was previously housed at the University of Puerto Rico at Mayaqüez. At NCSU, it joins the excellent Auchenorrhyncha collections of Zeno P. Metcalf, Wilhelm Wagner, and David A. Young.

José Andrés Ramos Alemar (1915-1989)

Treehopper Publications by José A. Ramos

Ramos, J. A. 1947a. The insects of Mona Island (West Indies). The Journal of Agriculture of the University of Puerto Rico 30: 1-74, Plates 1-2. [notes on Paradarnoides spp., including P. danforthi, n. sp.; based on Ramos 1945 (unpublished Master’s thesis, State College of Agriculture and Engineering of the University of North Carolina, Raleigh)]
Ramos, J. A. 1956a. Review of the auchenorhynchous [sic] Homoptera of Puerto Rico. Part III. Kinnaridae, Cercopidae, Membracidae and Cicadidae. Dissertation Abstracts 16(4): 62. [abstract of Ramos 1956 (unpublished dissertation, North Carolina State College, Raleigh); dissertation published as Ramos 1957a]
Ramos, J. A. 1957a. A review of the auchenorhynchous [sic] Homoptera of Puerto Rico. The Journal of Agriculture of the University of Puerto Rico 41(1):38-117. [with Spanish summary; Antillotolania, Deiroderes, Jibarita, and Spathenotus, n. gen., and 9 n. spp. of Membracidae]
Ramos, J. A. 1979a. Membracidae de la República Dominicana (Homoptera: Auchenorhyncha). Universidad de Puerto Rico (Mayagüez), Estación Experimental Agrícola Boletin 260: 1-71. [in Spanish with English summary; Monobeloides n. gen., and 17 n. spp. in various genera]
Ramos, J. A. 1988a. Zoogeography of the auchenorrhynchous Homoptera of the Greater Antilles (Hemiptera). pp. 61-70. In Liebherr, J. K. (ed.). Zoogeography of Caribbean Insects. Cornell University Press, Ithaca, New York. xii + 285 pp. [notes on Membracidae; lists 26 genera, p. 64]
Ramos, J. A. 1990a (dated 1989). New West Indian Membracidae (Homoptera: Auchenorrhyncha). Caribbean Journal of Science 25(3-4): 153-163. [notes on 10 n. spp. in 6 genera; publication dated 1989, but "Mailed February 1990”].

"Smithsonian Insider" Features Treehoppers
2017-08-31
Barrat, J. 2017a. Beautiful and bizarre, treehoppers suck sap and can spread disease. Smithsonian Insider (Animals, Earth Science, Plants, Q & A, Research News, Science & Nature Animals): 28 August 2017. [online publication]

See this popular article about treehoppers and USDA Systematic Entomology Laboratory scientist Stuart McKamey, a dedicated researcher and member of our Treehoppers Team.

New Collaborator and Links to Checklists with Treehopper Images
2017-09-12
To our lists of collaborators (see Treehoppers Acknowledgments and Archive of Past News), we add Todd F. Elliott (School of Environmental & Rural Sciences, University of New England, New South Wales, Australia).

Figs. 1-2. Todd took these wonderful photos of Anchon speciosum Capener in the Dja Biosphere Reserve in Cameroon; our team member Dawn Flynn identified the treehopper to species.

1 2 Images © 2017, by Todd F. Elliott

Speaking of treehopper images, among the growing number of online resources on treehoppers are the following illustrated checklists for Florida, North Carolina, Signapore, and Texas. Nicky Bay’s treehopper images from Signapore are exceptionally stunning.

About Ants and Treehoppers
2017-10-26
Many treehopper species benefit from interactions with ants that provide "house cleaning" services and sometimes protect the hoppers from predators in exchange for honeydew secreted by the hoppers. The images below show ants tending adults of Stilbophora luteimaculata (Fig. 1: © 2007, by Kelly Swing) and immatures of Bolbonota (Fig. 2: © 2013, by Kenji Nishida).

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To learn more about the relationships among ants and treehoppers, check out these blogs (with videos) at “Wild about Ants:” Ants Tending the Aster Hopper Publilia concava and Well-Guarded Treehoppers.

Phylogeny and Divergence Time Estimates within the Membracoidea
2017-11-15
Dietrich, C. H.; Allen, J. M.; Lemmon, A. R.; Lemmon, E. M.; Takiya, D. M.; Evangelista, O.; Walden, K. K. O.; Grady, P. G. S.; Johnson, K. P. 2017a. Anchored hybrid enrichment-based phylogenomics of leafhoppers and treehoppers (Hemiptera: Cicadomorpha: Membracoidea). Insect Systematics and Diversity 1(1): 57–72. [available online]

DNA from 388 loci and more than 99,000 aligned nucleotide positions were used to estimate phylogenetic relationships among 138 taxa representing all major lineages of the superfamily Membracoidea (leafhoppers and treehoppers). In agreement with several previous studies, this work indicates leafhoppers (family Cicadellidae) are paraphyletic with respect to treehoppers (families Aetalionidae, Melizoderidae, and Membracidae). Treehopper taxa represented in the analyses included 2 aetalionid genera (2 tribes), 1 melizoderid genus, and 39 membracid genera (32 tribes). Divergence time estimates suggest the lineage comprising the treehopper families arose during the Cretaceous. Within Membracidae, the authors found strong support for two major sister lineages: one with all treehoppers in which the pronotum extends posteriorly over the scutellum, but does not conceal it–and the other with nearly all treehoppers in which the pronotum entirely conceals the scutellum (but none with the scutellum concealed only in part).

Zoom in on Germar's Collection: Updated Link
2017-12-07
Zoomable digital images of the Ernst Friedrich Germar Collection are available online thanks to the curators of the Benedict Dybowski Zoological Museum (Lviv National University, Ukraine). For a number of years, this historically significant collection was thought to have been destroyed in World War II. A revised link to this collection is provided here and on our 4 March 2012 news item on Germar's Collection.