Recent Publications on Symbiont–Pathogen Diversity in Membracidae and the Phylogeny of Centrotinae
Salazar, M. M.; Pupo, M. T.; Brown, A. M. V. 2021a. Co-occurrence of viruses, plant pathogens, and symbionts in an underexplored hemipteran clade. Frontiers in Cellular and Infection Microbiology 11:715998. [Online publication].

Research on 16 treehopper species revealed that this insect group offers a rich resource for studying symbiont–plant pathogen interactions, including potential primary and secondary insect symbionts, potential plant pathogens, and other microbes and viruses.

Yu, R.; Feng, L.; Dietrich, C.H.; Yuan, X. 2022a. Characterization, comparison of four new mitogenomes of Centrotinae (Hemiptera: Membracidae) and phylogenetic implications supports new synonymy. Life 2022, 12, 61. [Online publication].

This work examines phylogenetic relationships within the subfamily Centrotinae based on mitochondrial genome data, incorporating newly sequenced mitogenomes of Anchon lineatus, Anchon yunnanensis, Gargara genistae, and Tricentrus longivalvulatus. The authors’ resulting phylogenetic analyses support Wallace and Deitz’s (2004a) placement of the tribes Tricentrini and Antialcidini as junior synonyms of Gargarini.

Recent Contributions on Heteronotus, Micrutalis, and Stylocentrus
Perger, R. 2021a. First reports of species-specific ant resemblance in heteronotine treehoppers (Hemiptera: Membracidae: Heteronotinae). Insecta Mundi: 0888:1-6. [Online publication].

This publication documents species-specific resemblance in females of the Bolivian treehopper Heteronotus fabulosus and workers of an associated giant turtle ant, Cephalotes atratus. Females of the treehopper (including multiple structural details of their shiny black, spiny pronota) appear to mimic an attending species of shiny black, spiny, turtle ant. Even though it does not sting, C. atratus, is a large, aggressive ant with intimidating spines, and thus an appropriately menacing model for such a mimetic relationship.

Wheeler, A. G., Jr.; Flynn, D. J. 2021a. First U.S. records of the mistletoe-associated Micrutalis discalis (Walker) (Hemiptera: Membracidae). Description of the male, and redescription of the female. Proceedings of the Entomological Society of Washington 123(3): 652-664.

The authors give the first US reports of M. discalis (Walker) from 5 counties in Arizona. This Micrutalis was originally described based on a single specimen lacking its abdomen (probably a female) from Veracruz, Mexico, in 1858. The present work provides a redescription of the female, along with the first descriptions of the male and the fifth instar based on adults and nymphs from desert mistletoe (Phoradendron californicum). Field observations suggest the species is bivoltine.

Updated Citation: Wang, Zhang, Shih, Dong & Yao (2020a) A new species of Membracidae (Hemiptera: Cicadomorpha: Membracoidea) from Dominican amber. On 2020-09-29, the Treehopper News listed this article based on the 2020 online version of the paper (6 August 2020). It is now also available in print form with the following date and pagination: Wang, H.; Zhang, X.; Shih, C.; Dong, R.; Yao, Y. 2021a. A new species of Membracidae (Hemiptera: Cicadomorpha: Membracoidea) from Dominican amber. Historical Biology 33(10), 2491-2495. [first published online in 2020]

A Case for the Commission: a Tribe of Treehoppers and a Tribe of Katydids Share Identical Names
Deitz, L. L.; Cryan, J. R. 2021a. Case 3846 –Microcentrini Deitz, 1975 (Insecta, Hemiptera, Membracidae): proposed emendation of spelling to Microcentrusini to remove homonymy with Microcentrini Brunner von Wattenwyl, 1878 (Insecta, Orthoptera, Phaneropteridae). Bulletin of Zoological Nomenclature 78: 96-98.

Deitz and Cryan referred this case to International Commission on Zoological Nomenclature to resolve the homonymy of the treehopper tribe Microcentrini Deitz, 1975 and the katydid tribe Microcentrini Brunner von Wattenwyl, 1878. The ICZN acknowledged the receipt of this application in April (Bulletin 78: 2), preceding its publication in August (78: 96-98). Comments on this case for publication in the Bulletin (subject to editing) may be sent to the Secretariat, International Commission on Zoological Nomenclature, c/o Lee Kong Chian Natural History Museum, 2 Conservatory Drive, Singapore 117377, Republic of Singapore (e-mail:

The Socially Distanced Treehopper Aggregation of 2021
The 29th Annual Treehopper Gathering was held 4-6 June 2021, at the Little Orleans Campground, Little Orleans, Maryland. The timing was fortunate because participants had time to become fully vaccinated against the COVID-19 virus before the gathering. Fourteen people and two "treehopper-curious" dogs attended this year (Figs. 1 and 2) and we had a lot of catching up to do.


Figs. 1-2. Group Photos. 1, fully masked and socially-distanced. 2, unmasked and less distanced (left to right in Fig. 2): Stuart McKamey, Lewis Deitz, Kelley Tilmon, Matthew Wallace, Andrew Short, Mark Rothschild, Eli Wyman, Hava Amsbury, Wil Winter, Dawn Flynn, Ashley Kennedy, Anthony Deczynski, and Jamie Zahniser, plus dogs "Shaggydog" (left) & "Delphine" (right) [Jason Owens is absent in Fig. 2, but is fourth from the left in Fig. 1]. (Figs. 1 & 2 © 2021, by Wil Winter and Ashley Kennedy).

Those attending were serenaded daily from dawn to dusk by periodical cicadas of Brood X at our campsite near Fifteenmile Creek that runs through the campground. The weather was pleasant with bright skies and butterflies. After the sunset on Saturday (5 June), with the aid of a portable generator, we watched "The Love Bugs" projected on a white sheet at our campsite. This charming documentary film features entomologists Lois and Charlie O’Brien, known widely for their work on planthoppers and weevils, respectively. Later, because the moon was in its waning crescent phase, treehoppers were especially abundant at various lights set up for night collecting. It was great to get together in person and catch up on happenings in the world of treehopperdom.

4 5 6
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Figs. 3-8. Sights around the campground. 3, scenic view of Fifteenmile Creek. 4, wild stonecrop, Sedum ternatum, near the creek. 5, Great Spangled Fritillary butterfly visiting a flowering shrub. 6, Mark Rothschild collecting treehoppers at light. 7, Wil Winter collecting with Andrew Short at the creek. 8, baby snapping turtle. (Copyrights: Figs. 3-6, © 2021, by Matt Wallace; Figs. 7-8, © 2021, by Ashley Kennedy).

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