Updates to Treehoppers Team Members and Collaborators
We welcome graduate student Camilo Flórez-Valencia as the newest member of the Treehoppers "Taxa Content Team." A native of Colombia, Camilo is currently working towards his Master’s degree with Brazilian treehopper researcher Olivia Evangelista. One of Camilo's photos, a Colombian Anobilia, is shown below.

New collaborators include entomologists René Limoges and Stéphane Le Tirant (Montreal Insectarium, Montréal, Québec, Canada). Two of René’s recent treehopper photographs from Mindo, Ecuador, are featured below. See our Acknowledgments and archive of past news (posting of 2015-05-12), for further listings of Team Members and Collaborators.

1, Anobilia nigra, adult with nymphs and ant; 2, Alchisme adult with nymphs; 3, Aconophora female with egg mass. Image copyrights: 1, © 2015, by Camilo Flórez-Valencia; 2 and 3 © 2015, by René Limoges.

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Treehopper Publications of E. P. Van Duzee and L. B. Woodruff
Most works on treehoppers by American entomologists Edward Payson Van Duzee (1861-1940) and Lewis Bartholomew Woodruff (1868-1925) are readily available online through the Biodiversity Heritage Library (BHL). Two of Woodruff’s papers (1923a, 1924a), not currently available at BHL, are accessible through the digital library JSTOR (see Options for Access). For detailed citations of the works by either author, enter his last name in the author search box of DrMetcalf. We note that "Van Duzee 1890a,” which seems to treat only extant leafhoppers, was apparently listed in Metcalf and Wade’s 1963 bibliography on treehoppers and fossil Auchenorrhyncha in error.

Hemipterist Edward P. Van Duzee described 3 treehopper genera (Idioderma, Tylocentrus, and Xantholobus), 43 species (36 now valid, including 2 that are also nominotypical subspecies) and 3 subspecies (2 now valid subspecies, 1 invalid). The California Academy of Sciences, where Van Duzee served as curator (1916-1940), holds his collection of Hemiptera (164,442 specimens). Van Duzee’s 1908a plates 1 and 2, below, show simple line drawings of representative North American treehoppers.

Van Duzee’s Treehopper Publications: Van Duzee 1889b; Van Duzee 1890c; Van Duzee 1892d; Van Duzee 1893b; Van Duzee 1894e; Van Duzee 1895e; Van Duzee 1905a; Van Duzee 1906a; Van Duzee 1907a; Van Duzee 1908a; Van Duzee 1908b; Van Duzee 1909a; Van Duzee 1909c; Van Duzee 1912a; Van Duzee 1912b; Van Duzee 1914a; Van Duzee 1914c; Van Duzee 1916a; Van Duzee 1916b; Van Duzee 1917a; Van Duzee 1917b; Van Duzee 1923a; Van Duzee 1925b.

Lewis B. Woodruff authored 18 treehopper species (16 now valid) and 2 subspecies (1 now elevated to species, 1 invalid), all belonging to the tribe Smiliini. See our news posting of 2012-09-14 [archive of past news], for further information on Woodruff, who at one time had a desk at the American Museum Natural History. The following plates are from Woodruff 1915a (pl. 4, figs. 1-5, now Atymna helena); fig. 6, now A. querci) and 1919a (pl. 23; Ophiderma spp. [Woodruff’s captions include some junior synonyms]).

Woodruff’s Treehopper Publications: Woodruff 1915a; Woodruff 1919a; Woodruff 1920a; Woodruff 1923a; Woodruff 1924a.

[Compiled by Matthew S. Wallace and Lewis L. Deitz, 9 November 2015]

Units Used to Measure Insects Prior to the Metric System
Be grateful that millimeters have largely replaced “lines” and “lunules” as units to measure treehoppers. By the way, a “lunule” is the white crescent-shaped area at the base of one’s fingernail--lunules vary in size and may even be absent in some individuals. More frequently, early entomological workers used “lines” as a unit of measurement, but the definition of a line varied among the English, French (ligne), and Germans (linie), and, in some instances, the definition varied even within a country. The Website includes a brief overview of early entomological measurements. To convert some of the more commonly used "lines" to millimeters, see “Notes on the line as a unit of measurement,” p. 11, from von Hayek, 1973, Bulletin of the British Museum (Natural History), Entomology Supplement 20: 1-309.

Guide to Treehopper Workers of the 1700s and Their Publications
This guide provides access to virtual copies of most of the literature on treehoppers by scholars of the 1700s, including their further works into the 1800s. One exception is Pierre A. Latreille, whose 1796a work is listed here, but not his 24 additional references on treehoppers published during the 1800s. Also excluded here are workers, such as H. R. Scheller, whose first publication appeared after 1799. The earliest work listed here is Linnaeus’s (1746a: p. 202) description of his species number 641, subsequently named Centrotus cornutus Linnaeus, 1758. Owing to modern technology and awe-inspiring international collaboration, this collection of rare and priceless publications is at your fingertips.

Early contributions on treehoppers were sometimes published over a period of two or more years and subsequently bound into a single volume with a title page and introductory materials. Because of the principle of priority among taxonomic names, it is essential to establish the actual date of publication for each section of such works that describe new taxa (the section date may differ from that on the title page). Furthermore, the sequence of sections and placement of plates may vary among various bound copies of a work. In the absence of photography, illustrations were frequently produced from copper engravings (plates), and some were individually hand-colored, each plate being a unique work of art.

Selected early natural history plates with figures of treehoppers: 1, Coquebert 1799a: Pl. 8, Fig. 1; 2 , Coquebert 1799a: Pl. 9, Fig. 5; 3, Coquebert 1801a: Pl. 18, Figs. 1-10; 4, Coquebert 1804a: Pl. 21, Fig. 5; 5, Houttuyn 1766a: Pl. 81, Fig. 2; 6, Linnaeus 1761a: Pl. 2, Fig. 279; 7, Panzer 1798a: Pl. 50.19, Membracis cornutus; 8, Panzer 1798a: Pl. 50.20, Membracis genistae; 9, Donovan 1794a: Pl. 83; 10, Donovan 1820d: Hemiptera Pl. 3, C. spinosa, C. taurus; 11, Stoll 1788a: Pl. [28], Figs. 163, A, B, 165, D, 166, E, 169, H; 12, Sulzer 1761a: Pl. 10, Fig. 63; 13, Sulzer 1776a: Pl. 9, Fig. 6.

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Z. P. Metcalf took care in establishing publication dates, and we follow his dates unless convincing evidence has come to our attention indicating a needed correction. In this regard, during the compilation of the present guide, we discovered Index Novus Litteraturae Entomologicae, an updated bibliography of the literature on entomology from the beginning until 1863, which appears to be a useful resource for verifying publication dates. To generate full citations for nearly all works published on Auchenorrhyncha during the 1700s, enter "17??" in the date search box of DrMetcalf.

The names of early authors often vary in spelling depending upon the language of the publication, thus variants are given in square brackets in the following list of Treehopper Workers of the 1700s: James [Jacque] Barbut; Johann Beckmann [Iohanne Beckmanno]; Nikolaus Joseph [Nicolaus Josephus] Brahm; Antoine Jean [Antoine-Jean, Antonius-Joannes, Antoine-Jean-Joseph, Jean Antoine] Coquebert [Coquebert de Montbret]; Charles [Caroli] De Geer; Charles [Carolo] Joseph De Villers [de Villers]; Edward Donovan; Johann [Iohannes, Jean, Johan, Johann, Johannes] Christian [Christianus, Chrtien] Fabricius; Johann Reinhold Forster [Joanne Reinholdo Forstero]; Antoine François [Francois, Francoise] Fourcroy [de Fourcroy]; Johann Kaspar [Caspar] Füssli [Fuessli, Fuesslins, Fuessly]; Étienne [Etienne] Louis Geoffroy; Michel Esprit Giorna; Johann Friedrich Gmelin; Johann August Ephraim Goeze; Laurens [Laurentius] Theodorus Gronovius; P. Höslin; Martin [Martinus] Houttuyn; Pierre André [Andre] Latreille; Anton August Heinrich Lichtenstein; Carolus Linnaeus [Carl von Linné]; Christian Friedrich Ludwig; Maria [María, Marie, Anna Maria] Sybilla [Sybille] Merian [de Merian]; Otto Frederik [Friedrich] Müller [Mueller]; Philipp Ludwig Statius Müller [Mueller]; Guillaume Antoine Olivier; Peter Simon Pallas; Georg Wolfgang Franz Panzer; Vincenzo Petagna; Erich [Erik] Pontoppidan; Anders Jåhan [Jahan] Retzius; Johann Jacob Römer [Roemer]; Pietro Rossi; Jacob [Jakob] Christian Schäffer [Schaeffer]; Franz von Paula [de Paula] von Schrank [Schrank]; Johann Christian Daniel von Schreber; Giovanni [Johannes, Ioannis] Antonio [Antonii] Scopoli; Caspar Stoll; Johann [Johan] Heinrich Sulzer; Carl Peter [Caroli Petri, Carolo Petro] Thunberg [Thvnberg]; Friedrich [Frederico] Weber; Thomas Pattinson Yeats; Johann Jacob [Iohannes I.] Zschach [Zschachii] and Nathanael Gottfried Leske [Leskeanum].

Publications on Treehoppers by Workers of the 1700s: Barbut 1783a, pl. 9 ; Beckmann 1772a; Brahm 1790a; Coquebert 1799a, pls. 8-9; Coquebert 1801a, pl. 18; Coquebert 1804a, pl. 21; De Geer 1773a [see pp. 158-328 and pl. 11, Fig. 23, image incomplete]; De Geer 1778a, [see pp. 158-228, Cigales], [plate 45]; De Villers 1789a; Donovan 1794a, pl. 83; Donovan 1820a, [see Cicada]; Donovan 1820d, Hemiptera pl. 3, C. spinosa, C. taurus; Fabricius 1775a; Fabricius 1777a; Fabricius 1781a; Fabricius 1787a; Fabricius 1794a; Fabricius 1798a; Fabricius 1803a; Forster 1771a; Fourcroy 1785a; Füssli 1775a; Geoffroy 1762a, pl. 9 ; Geoffroy 1764a; Geoffroy 1799a; Giorna 1791a; Gmelin 1789a; Goeze 1778a; Goeze 1780a [needed]; Gronovius 1762a [needed], [see ICZN Opinion 261: rejected for nomenclatural purposes]; Höslin 1782a [needed]; Houttuyn 1766a, pl. 81; Latreille 1796a [24 treehopper publications in the 1800s are not listed here]; Lichtenstein 1796a, [see ICZN (1995) Opinion 1820, Bulletin Zoological Nomenclature 51: 283-285]; Linnaeus 1746a [Laurentii Slavii printing], [Lugundi Batavorum printing]; Linnaeus 1758a; Linnaeus 1761a, pl. 2; Linnaeus 1764a; Linnaeus 1767a [subsequent reprints of 1758a not listed here]; Ludwig 1799a; Merian 1771c; Müller, O. 1776a; Müller, P. L. S. 1774a [needed]; Olivier 1789a; Olivier 1792a; Olivier 1797a, pl. 109; Pallas 1766a, pl. 14; Pallas 1772a, pl. 1; Panzer 1798a, pls. 50.19-50.20; Panzer 1802a [needed]; Panzer 1804a; Petagna 1787a; Petagna 1792a, pl. 9; Petagna 1808a [needed]; Petagna 1820a [needed]; Pontoppidan 1762a [see p. 680, Hemiptera, and pl. 29 Cicada cornuta]; Retzius 1783a; Römer 1789a, pl. 9; Rossi 1792a, [see pp. 212-220, Rhyngota]; Rossi 1794a, [see p. 50, Ryngota]; Rossi 1807a, [see pp. 341-352, Rhyngota]; Schaeffer 1767a, pl. 96; Scheller 1869, [plates]; von Schrank 1781a; von Schrank 1801a; von Schreber 1759a; Scopoli 1763a; Scopoli 1777a; Stoll 1781a; Stoll 1788a, pls. 1-2, pls. 5-6, pl. 8, pls. 10-11, pls. 15-[19], pl. 21, pl. [28]; Stoll 1792a [German translation of 1788a]; Sulzer 1761a, [see pp. 94-98, 24-25 and pl. 10]; Sulzer 1776a, pl. 9; Thunberg 1789a; Thunberg 1791a; Thunberg 1822a [needed]; Weber 1795a, [see pp. 144-148, Rhyngota; Weber 1801a, [see pp. 112-115, Rhyngota]; Yeats 1773a; Zschach and Leske 1788a, [see pp. 115-117, Ryngota].

Please contact Lewis Deitz if you know of other works mentioning treehoppers printed prior to 1800 or if you can provide links to online copies of works marked here as “needed.”

archive of past news