Join the 29th Annual Treehopper Gathering: a Socially Distanced Aggregation
The 2021 Treehopper Gathering is arranged for 4-6 June 2021 at the Little Orleans Campground, Little Orleans, Maryland. This event provides a beautiful, casual setting to meet and collect with others passionate about treehoppers and related insects in the 222-acre campground, located in the Upper Potomac River valley of western Maryland. At least 23 treehopper genera (56 species) have been collected in this area.

To reserve a campsite with our group or to obtain further information, contact Charles Bartlett. Advanced registration with Charles is essential to assure that you have a campsite. Because of the pandemic, we strongly recommend that all attendees be fully vaccinated.

Participants are expected to provide their own tents, food, and camping gear, and to share in the cost of the group campground registration. Note that this year a socially distanced aggregation time is planned for Saturday evening, rather than the usual shared group meal. We hope to be able to view the documentary film The Love Bugs.

Bring your collecting vials, insect nets, unidentified specimens, cameras, bathing suits, a folding chair, and your favorite t-shirts. If you fear cicadas, you may also wish to bring a wearable pod to protect you from Brood X of 17-year cicadas. LOL!

New Online Publication: Third Version of "Guide Illustré des Membracides de Guyane" by Jérémie Lapéze
Lapèze, J. 2021a. Guide Illustré des Membracides de Guyane. Version 3. 142 digital pp. Available online: [in French]

This beautiful and useful illustrated guide to the treehoppers of French Guiana includes, in the words of Jérémie Lapèze, “more species, more photos, more information.”

David Stephan and His Telamona
We are saddened to report the recent passing of David Leigh Stephan (1946-2021), for whom the Nearctic treehopper Telamona stephani is named (Wallace 2018a). Dave was an extraordinary naturalist, with an amazing knowledge of the North Carolina flora and fauna. For his entire career at North Carolina State University, he served as the chief insect identification specialist for the NC Plant Disease and Insect Clinic. Much admired as a naturalist and conservationist, Dave will be greatly missed.

It is most fitting that the beautiful treehopper shown below will forever remind us of our dear friend and colleague. David collected both the holotype (Fig. 1) and allotype (Fig. 2) at lights in Bladen County, North Carolina, however, this sexually dimorphic species occurs also in Delaware, Louisiana, Maryland, Pennsylvania, and Virginia (Wallace 2018a). Males differ from other species of Telamona in having a characteristic mottled coloration, a dark brown head and metopidium, and very lowly elevated pronotal projection. Little is known of the biology of this treehopper because most specimens were taken at lights.

1 2

Figs. 1 and 2, Telamona stephani. 1, holotype, male. 2, allotype, female. Both specimens are at the NCSU Insect Collection. Photos © 2021, by Matthew A. Bertone.

Wallace, M. S. 2018a. A new species of Telamonini from the eastern United States and the taxonomic limits of Telamona salvini Distant (Hemiptera: Membracidae: Smiliinae). Proceedings of the Entomological Society of Washington 120(3):605-615.

Publications on Treehoppers by Adachi et al. 2000a and Flynn and Gálvez 2021a
Adachi, H.; Matsuda, K.; Nishida, K.; Hanson, P.; Kondo, S.; Gotoh, H. 2000a. Structure and development of the complex helmet of treehoppers (Insecta: Hemiptera: Membracidae). Zoological Letters 6, 3 (2020). Available online:

This research on the morphogenesis of prothoracic structures in Antianthe expansa was based on micro-CT, scanning electron microscopy, and paraffin sections. It presents excellent illustrations, including supplemental videos of eclosion in the genera Antianthe, Bocydium, Cladonota, Poppea, and Umbonia.

Flynn, D. J.: Gálvez, J. E. B. 2021a. A new treehopper species of Cladonota (Stål) (Hemiptera: Membracidae: Membracinae: Hypsoprorini) from Mexico. Zootaxa 4948(4): 569-574.

Cladonota (Lecythifera) crucifixa Gálvez and Flynn in Flynn and Gálvez, n. sp., is illustrated and described from Puebla, Mexico. Its host plant is Euphorbia schlechtendalii, however, based on the manuscript, the host species name seems to be misspelled on the data labels. Both sexes of this new treehopper (Fig. l) resemble females of C. hoffmanni.

Figure 1. Cladonota (Lecythifera) crucifixa, female. Image © 2021, by Juan Enrique Benítez Gálvez (Puebla, Puebla, Mexico). We thank Enrique Gálvez for sharing this image as a new collaborator.

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